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1972:302 [catalog listing as subspecies]; Martínez 1976:327 [distribution]. Platycoelia pomacea pomacea, Martínez 1976:327 [distribution]. TYPE SPECIMENS. Platycoelia pomacea Erichson lectotype female at ZMHB labeled a) “11465” (typeface), b) “Type” (orange label, typeface), c) “Pomacea (N.) Er.* Peru. Pavon.” (green label, handwritten), d) “PLATYCOE- LIA POMACEA ERICHSON F DET:A.B.T. SMITH 2001 LECTOTYPE” (red label, handwritten and typeface). Lectotype here designated. See Methods and Materials section for a statement of taxonomic purpose. Erichson (1847) did not state how many specimens were in the type series. The existence and location of paralectotypes are unknown. Type locality: Perú. Platycoelia boliviensis Blanchard lectotype male at MNHN labeled a) “6436 34” (round, green label, handwritten on underside), b) “P. boliviensis. Cat. Mus. Santa-Cruz. (Bolivie). M. d’Orbigny.” (green label, handwritten), c) “PLATYCOELIA BOLIVIENSIS BLANCHARD M LECTOTYPE A. B. T. SMITH 2002” (red label, handwritten and typeface), d) “Platycoelia pomacea Erichson, 1847 M Det:A.B.T.Smith 2002” (typeface). Lectotype here designated. See Methods and Materials section for a statement of taxonomic purpose. Blanchard (1851) did not indicate how many specimens were in the type series (although the range of lengths indicates more than one). The location of the paralectotype(s) is unknown. Some of d’Orbigny’s handwritten notebooks on the specimens in his collection are at the MNHN library. The entry for the type series of this name states “Areoda 1437. Melolontha. Viz dans les grande forêt, sur les tigre de arbuste. Je en nocturne ou au moisse crépusculaire.” Ohaus (1904b) misapplied the name P. boliviensis to P. peruviana. He did examine the lectotype of P. boliviensis during his visit to Paris during 1903, but he did not carefully examine it or dissect the genitalia. The lectotype of P. boliviensis is, without a doubt, P. pomacea. Therefore I am placing these names in synonymy. Type locality: Santa Cruz, Bolivia. NEW SYNONYMY. MONOGRAPHIC REVISION OF PLATYCOELIA 67 Platycoelia pomacea brasiliensis Ohaus lectotype male at ZMHB labeled a) “Planalto d. Parecis” (typeface), b) “Pl. pomacea brasiliensis Type Ohs.” (orange label, handwritten), c) “PLATYCOELIA POMACEA BRASILIENSIS OHAUS LECTOTYPE A.B.T.SMITH 2001” (red label, handwritten and typeface), d) “Platycoelia pomacea Erichson, 1847 M Det:A.B.T.Smith 2002” (typeface). Lectotype here designated. See Methods and Materials section for a statement of taxonomic purpose. Three male and one female paralectotypes at MNHN labeled a) “Jatahy Etat de Goyaz Ch. Pujol 1895-96” (typeface), b) “Ohaus determ. Pl. pomacea Er. v. brasiliensis Ohs. [M/F]” (typeface and handwritten), c) “Dr Ohaus Vidit 1903.” (typeface), d) “PLATYCOELIA POMACEA BRASILIEN- SIS OHAUS PARALECTOTYPE A.B.T. SMITH 2001” (yellow label, handwritten and typeface), e) “Platycoelia pomacea Erichson, 1847 [M/F] Det:A.B.T.Smith 2002” (typeface). Four female paralectotypes at MNHN labeled as previous paralectotype omitting label b. One male paralectotype at ZMHB labeled a) “Jatahy (GOYAZ)” (typeface), b) “M” (typeface), c) “Pl. pomacea brasiliensis Cotype Ohs.” (orange label, handwritten), d) “PLATY- COELIA POMACEA BRASILIENSIS OHAUS PARALECTOTYPE A.B.T.SMITH 2001” (yellow label, handwritten and typeface), e) “Platycoelia pomacea Erichson, 1847 M Det:A.B.T.Smith 2002” (typeface). One female paralectotype at ZMHB labeled as previous paralectotype except label b) “M ” (typeface). One male paralectotype at ZMHB labeled a) “Jatahy Goyaz” (typeface), b) “Pl. pomacea brasiliensis Cotype Ohs.” (orange label, handwritten), c) “PLATYCOELIA POMACEA BRASILIENSIS OHAUS PARA- LECTOTYPE A.B.T.SMITH 2001” (yellow label, handwritten and typeface), d) “Platycoelia pomacea Erichson, 1847 M Det: A.B.T. Smith 2002” (typeface). One female paralectotype at ZMHB labeled a) “Jatahy Goyaz” (typeface), b) “Sept-Nov. 97.” (handwritten), c) “F” (typeface), d) “Pl. pomacea brasiliensis Cotype Ohs.” (orange label, handwritten), e) “PLATY- COELIA POMACEA BRASILIENSIS OHAUS PARALECTOTYPE A.B.T.SMITH 2001” (yellow label, handwritten and type-
68 BULLETIN OF THE UNIVERSITYOF NEBRASKA STATE MUSEUM face), f) “Platycoelia pomacea Erichson, 1847 F Det: A. B.T. Smith 2002” (typeface). One female paralectotype at ZMHB labeled a) “Miranda, XI.94.-II. 95. (Andeer)” (typeface), b) “F” (typeface), c) “Pl. pomacea brasiliensis Cotype Ohs.” (orange label, handwritten), d) “PLATYCOELIA POMACEA BRASILIENSIS OHAUS PARALECTOTYPE A.B.T.SMITH 2001” (yellow label, handwritten and typeface), e) “Platycoelia pomacea Erichson, 1847 F Det: A.B.T. Smith 2002” (typeface). The existence and location of other paralectotypes are unknown. There is no character-based justification for maintaining this subspecies as separate from P. pomacea, therefore I am placing the two names in synonymy. Type locality: Planalto do Panecis, Mato Grosso, Brazil. NEW SYNONYMY. DESCRIPTION. Male (n=68). Length 21.2- 27.1 mm, width 12.9-17.1 mm. Color light green with thick yellow elytral suture; yellow tarsi, mouthparts; yellow or brown around some margins, sutures. Body ovate, strongly convex. Head: Dorsal surface glabrous. Frons and clypeus sparsely punctate, punctures moderate. Frons not depressed. Frontoclypeal suture complete. Clypeal apex rounded. Eyes separated by approximately 3.5 transverse eye-widths. Labrum sparsely punctate, with moderately large, setose punctures, setae tawny. Apex of labrum with triangular medial tooth, apex of tooth truncate, extending past apex of mentum. Mandibular scissorial region with strong apical tooth, weak medial tooth; molar region with well-developed lamellae. Maxilla with 3 apical, cup-like teeth. Apex of mentum with triangular medial tooth curved into oral cavity. Antenna 10-segmented; club slightly longer than other segments combined, subequal to length of frons. Pronotum: Surface glabrous, sparsely punctate, with moderate punctures. Marginal bead weak laterally, absent apically and basally. Elytron: Surface glabrous; longitudinal striae weakly punctate, punctures moderate with dark coloration; intervals sparsely punctate to impunctate. Suture with distinct apical spine. Pygidium: Width approximately 3.0 times length medially. Surface rugose, weakly convex, glabrous except near apex; setae short, tawny. Venter: Thorax glabrous. Mesothoracic process strongly produced, projecting anteriorly to protrochanter; apex rounded; shape triangular-elongate, dorsoventrally flattened. Abdomen glabrous. Apical spiracles not extruding. Legs: Protibia with 2 teeth in apical half; apical tooth longer. Mesotibia and metatibia widest medially. Protarsomeres 2- 4 wider than long, cup-shaped. Protarsomere 4 with internoapical stridulatory ridge. Protarsomere 5 with elongated internomedial tooth bearing stridulatory ridge. Mesotarsomere and metatarsomere 5 without internal tooth. Unguitractor plate cylindrical, with 1 apical and 1 subapical seta. Modified protarsal claw length less than protarsomere 5, thickened and elongate when compared with other claw, dorsolaterally flattened, apex unevenly bifurcate. Modified mesotarsal and metatarsal claws elongated with ventral tooth, not thickened. Male Genitalia: Phallobase 1.1 times longer than length of parameres. Parameres smooth, with weak apical bulb. Female (n=57). Length 23.0-29.0 mm, width 14.9-16.5 mm. As male except in the following respects. Head: Antennal club approximately equal to segments 2-7. Legs: Protarsomere 4 without internoapical stridulatory ridge. Protarsomere 5 with internomedial tooth absent. Modified protarsal claw with ventral tooth, not thickened. DIAGNOSIS. This species is distinguished from all other species in the genus Platycoelia by the following combination of characters: frontoclypeal suture complete; apex of the clypeus not reflexed; labrum with a well-developed, truncate apical tooth strongly overlapping the mentum; mentum with a distinct tooth curved into the oral cavity; antenna 10segmented; protibia with 2 prominent teeth; modified protarsal claw of male diagonally flattened; elytral suture distinctly yellow, with the apex forming an acute spine; elytron without yellow, longitudinal lines; elytral apices not capable of completely covering the pygidium; metasternum glabrous; metasternum and abdominal sternites uniformly
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Bulletin of the University of Nebra
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C O N T E N T S Abstract...........
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LIST OF FIGURES Fig. 1. Platycoelia
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Bulletin of the University of Nebra
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Members of the genus Platycoelia De
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alone to divide up the rutelines (a
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eing the most recent authority on t
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els Amongst the Great Andes of the
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tion. Zischka was a German who live
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Puncture size: large = 0.17 mm or l
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26. Pronotal surface sparsely punct
Page 24 and 25: Fig. 11. Platycoelia alternans male
Page 26 and 27: MONOGRAPHIC REVISION OF PLATYCOELIA
Page 28 and 29: Fig. 33. Platycoelia rufosignata me
Page 30 and 31: CNCI Canadian National Collection o
Page 32 and 33: subspecies AND varieties in the sam
Page 34 and 35: PHYLOGENETIC PATTERNS AND BIOGEOGRA
Page 36 and 37: metaunguitractor plate usually with
Page 40 and 41: larvae, and Paucar-Cabrera and Smit
Page 50 and 51: Erichson (1847) did not state how m
Page 52 and 53: species]; Ohaus 1918:178 [catalog l
Page 58 and 59: Platycoelia marginata and Platycoel
Page 60 and 61: tarsomere and metatarsomere 5 with
Page 62 and 63: 5. Platycoelia meridensis new speci
Page 64 and 65: in apical half. Protarsomere 4 with
Page 68 and 69: 7. Platycoelia prasina Erichson, 18
Page 70 and 71: written and typeface), e) “PLATYC
Page 72 and 73: mentum. Apex of mentum with triangu
Page 76 and 77: green to yellow; parameres with the
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Page 84 and 85: Mnisnech,” “Dr Ohaus Vidit 1903
Page 86 and 87: VARIATION. Male (n=19). Length 19.2
Page 88 and 89: “Platycoelia forcipalis cotype Oh
Page 90 and 91: males with internomedial stridulato
Page 94 and 95: tures moderate with dark coloration
Page 96 and 97: gins. Ventrally with dark brown met
Page 98 and 99: 19. Platycoelia flavostriata (Latre
Page 102 and 103: 20. Platycoelia nigrosternalis Ohau
Page 104 and 105: LOCALITY DATA. 66 specimens examine
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MONOGRAPHIC REVISION OF PLATYCOELIA
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Calloodes nitidissimus Lea, 1919 (C
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