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metaunguitractor plate usually with 1 apical,<br />

1 subapical seta (characters 64, 70); and each<br />

paramere apically rounded, usually not expanded,<br />

and usually without a setose patch<br />

(characters 74, 75, 78). This clade was part <strong>of</strong><br />

the “flavostriata group” <strong>of</strong> Ohaus (1904b) and<br />

was split into various groups by Machatschke<br />

(1965). The clade is found in the Andes Mountains<br />

from Venezuela to Bolivia. Members <strong>of</strong><br />

this clade generally occur in the Montane<br />

Wet Forest regions on the eastern slope <strong>of</strong> the<br />

Andes Mountains but several groups also occur<br />

in the high elevation Páramo and Puna<br />

regions. The Páramo region is a patchwork <strong>of</strong><br />

MONOGRAPHIC REVISION OF PLATYCOELIA 29<br />

moist habitats from Venezuela to northern<br />

Perú above continuous treeline. The Puna region<br />

is an extensive area in Perú, Bolivia,<br />

and northern Argentina above continuous<br />

treeline but much drier than the Páramo region.<br />

Several species in this clade have adaptations<br />

and modifications that presumably<br />

allow for survival in harsh, montane conditions<br />

present in the Páramo and Puna regions.<br />

The increasing elevation <strong>of</strong> the Andes Mountains<br />

throughout the Tertiary resulted in the<br />

Páramo and Puna regions and probably facilitated<br />

speciation within this clade <strong>of</strong><br />

Platycoelia.<br />

Fig. 38. Phylogram <strong>of</strong> the species in the genus Platycoelia. This is one <strong>of</strong> the 9 equally parsimonious trees<br />

found after successive approximation weighting. The treelength = 157.5; consistency index = 0.401; retention<br />

index 0.724; rescaled consistency index = 0.290.

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