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MONOGRAPHIC REVISION OF PLATYCOELIA 107 ano 13 May 1994 BL&MV DCCarlson/ FTHovore.” Thirteen male and six female paratypes at EGRC, two male paratypes at PKLC, two female paratypes at LACM, one male and one female paratypes at ABTS, one male and one female paratypes at ZMHB, and one male and one female paratypes at MLJC labeled “PANAMA: Cocle, Cerro Gaital, VI- 10-12-1985, E.Riley&D.Rider.” One male and one female paratype at DJCC and one male and one female paratype at ABTS labeled “PANAMA: Coclé Cerro Gaital 850m 13-20 vii 96 Curoe.” Two male and three female paratypes at DJCC labeled “Panama Cocle Cerro Gaital 11-V-94 Curoe, Monzon.” Two male paratypes at DJCC labeled “Panama Cocle prov. El Valle 850m 23 VIII 98 D. Curoe col.” One male paratype at SEMC labeled “PANAMA: Colon Prov (Santa Rita Ridge) 30 km SE Colon 13 May 1981 Robert W. Brooks.” Three male and one female paratypes at INBC and one male paratype at BCRC labeled “COSTA RICA, Prov. Alajuela, R. B. San Ramón. 800m. MAR 1997. G. Carballo. I_N_245100_472100 #46303” and have database labels numbering from INBIO CR1002 461158 to INBIO CR1002 461162. Five male paratypes at INBC, one male paratype at ABTS, one male paratype at UNSM, and one female paratype at BCRC labeled “R. San Lorencito, 900 m, R. F. San Ramón, 5 km N de Colonia Palmareña, Prov. Alaju., COSTA RICA. 13-18 Jun 1993. I Curso Scarabaeidae. L-N-244500, 470700” and have database labels numbering from INBIO CR1001 392210 to INBIO CR1001 392212, INBIO CR1001 364826, INBIO CR1001 364798, and from INBIO CR1001 392225 to INBIO CR1001 392227. One male paratype at INBC labeled “Sector Colonia Palmareña, Prov. Alaju, COSTA RICA. 700m. 3-22 ABR 1995. E. Fletes, L_N_245900_475900 #4770” and has the database label number INBIO CR1002 248190. One male paratype at INBC labeled “Sector Colonia Palmarena, Prov. Alaju, COSTA RICA. 700m. ABR 1995. G. Carballo, L_N_245900_475900 #5455” and has the database label number INBIO CR1002 213454. One male and one female paratypes at INBC labeled “COSTA RICA, Prov. Alajuela, Sector Colonia Palmareña, 9 Km SO. De Baja Rodriguez. 700m. MAY 1997. G. Carballo. L_N_245900_475900 #46301” and have the database label numbering from INBIO CR1002 541295 to INBIO CR1002 541296. One male paratype at ABTS labeled “COSTA RICA: Prov. Alajuela; Arenal Volcano (Observatory Lodge). 10°26’39"N, 84°42’62"W, 7- 17Apr. 1991, M. Spencer.” One male paratype at HAHC labeled “Limon Costa Rica Guayacan 8.IX.1972 V. O. Becker col.” One male paratype at INBC labeled “R. San Lorenzo, 1050m, R.F. Cord. Guanacaste (Tenorio) Prov. Guan. COSTA RICA. C. Alvarado, Jun 1991, L-N-287800,427600” and has the database label number INBIO CR1000 668623. One male and two female paratypes at INBC labeled “C. R.–San José P.N.B.C. Est. Carrillo Abril 1985 700m A. M. Chacón” and has the database label numbering from INBIO CR1002 519146 to INBIO CR1002 519148. One male paratype at CNCI labeled “COSTA RICA B. Carrillo N. P. 84°07’W;10°10’N 10.IV.85; 500m. H. Goulet- L. Masner.” One male paratype at USNM labeled “Rio Tacarcuna Darien Prov. Panama 1900 ft. Jul 63 W. P. Murdoch.” One female paratype at FSCA, labeled “PANAMA, PANAMA PR. ±10 KM N EL LLANO, 1400’ MAY 28 – JUNE 3, 1984 E. GIESBERT, COLL.” One female paratype at DJCC labeled “PANAMA: Coclé Cerro Gaital 850 m 15-23 vi 96 Curoe.” One female paratype from DJCC labeled “Panama, Pan Cerro Jefe 850m Altos de Capira 5 V 97 Curoe.” One female paratype at SEMC labeled “PANAMA 800m Panama Prov. Cerro Compana May 8, 1973” and “P D Ashlock Collector.” Two female paratypes at UNSM labeled “PANAMA: Panama Cerro Azul V-30-1986 B.Ratcliffe & party.” One female paratype at UNSM labeled “PANAMA, Chiriqui pr. 10 km NE FORTUNA DAM May 15-17, 1993 E. Giesbert, Coll.” and “Donated by Ed Giesbert 1997.” One female paratype at UNSM labeled “PANAMA: Bocas d. Toro 2 mi. N of Divide on hwy. To Chiriqui Grande VI-1- 1986 B.C. Ratcliffe & party.” One female paratype at ABTS labeled “COSTA RICA, Prov. Alajuela, Puesto Quebradón. P.N. Volcán tenorio 300m. ABR 1997. G. Rodriguez. L_N_292050_436700 #46262” and has the database label number INBIO CR1002
108 BULLETIN OF THE UNIVERSITYOF NEBRASKA STATE MUSEUM 553783. One male paratype at DCHC and one female paratype at UNSM labeled “COSTA RICA: Limón Guayacan 500 m 23-V-2001.” One female paratype at DCHC labeled “PANAMÁ: Coclé El Valle 850 m V-2000 D.Curoe.” Two male paratypes at FSCA labeled “PANAMA: Cocle Altos de Campana mv + bl. 13 May 1996 R. Turnbow” and “Platycoelia humeralis Bates det. R. Turnbow.” One male paratype at FSCA labeled “PANAMA, PANAMA PR. ±10 KM N EL LLANO, 1400’ MAY 28-JUNE 3, 1984 E. GIESBERT, COLL.” Two male paratypes at FSCA labeled “PANAMA, Panama pr. 7-10km N El Llano April 21-30, 1995 E. Giesbert, coll.” One male paratype at FSCA labeled “PANAMA, Panamá pr. 8-10km N El Llano May 24-June 2, 1992 E. Giesbert, coll.” One male paratype at FSCA labeled “PANAMA, Panamá pr. 8- 10km N El Llano Apr 26-May 4, 1992 E. Giesbert, coll.” One male and one female paratypes at ABTS labeled “PANAMA, Panamá pr. 13km N El Llano April 21-30, 1995 E. Giesbert, coll.” One male paratype at RFMC labeled “PANAMA Pnma Prv K 8-13 El Llano-Carti Rd 10-13 May 1996 Wappes Huether & Morris.” One male paratype at RFMC labeled “PAN:Panama Prov. 7-11 km El Llano/Carti Rd. 17-18/V/1999 Morris/ Wappes” and “Platycoelia humeralis Bates det. R. Morris.” Type locality: 10 km north of El Llano, Panamá, Panamá. HOLOTYPE. Male: length 26.9 mm, width 16.8 mm. Color light green with yellow head margin near eye, elytron, pronotum, scutellum margins, elytral stria 1. Body ovate, strongly convex. Head: Dorsal surface glabrous. Frons densely punctate, clypeus rugopunctate, punctures small or moderate. Frons not depressed. Frontoclypeal suture incomplete, absent medially. Clypeus trapezoidal. Eyes separated by approximately 4.5 transverse eye-widths. Labrum sparsely punctate, with moderately large, setose punctures, setae tawny. Apex of labrum with triangular, medial tooth; apex of tooth broadly truncate, extending past apex of mentum (as in Fig. 2). Apex of mentum with triangular medial tooth curved into oral cavity. Antenna 10-segmented; club slightly shorter than other segments combined, slightly shorter than length of frons. Pronotum: Surface glabrous, densely punctate, with small or moderate punctures. Marginal bead weak apicolaterally, absent elsewhere. Elytron: Surface glabrous; longitudinal striae weakly punctate, punctures moderate with dark coloration; intervals densely punctate, punctures small. Suture angled apically, without spine. Pygidium: Width approximately 3.0 times length medially. Surface weakly convex, moderately to densely punctate; punctures moderate, glabrous except near apex; setae short, tawny. Venter: Thorax moderately setose, setae white. Mesothoracic process strongly produced, projecting anteriorly past protrochanter; shape conical-elongate, apex rounded, dorsoventrally flattened (as in Fig. 8). Abdomen glabrous. Apical spiracles not extruding. Legs: Protibia with 3 subequal in size teeth in apical half; apical tooth longer. Mesotibia and metatibia widest medially. Protarsomeres 2-4 wider than long, cupshaped. Protarsomere 4 with internoapical stridulatory ridge. Protarsomere 5 with internomedial stridulatory tooth. Mesotarsomere and metatarsomere 5 without internal tooth. Prounguitractor plate cylindrical, with 2 subapical setae. Mesounguitractor plate and metaunguitractor plate cylindrical, with 2 apical setae. Modified protarsal claw with length approximately equal to protarsomere 5, thickened and elongate when compared with other claw, laterally flattened, apex unevenly bifurcate with 1 bifurcation twisted beneath the other in ventral view (as in Fig. 13). Modified mesotarsal and metatarsal claws elongated with ventral tooth, not thickened. ALLOTYPE. Female: length 29.5 mm, width 17.0 mm. As male except in the following respects. Head: Antennal club shorter than segments 2-7. Legs: Protarsomere 4 without internoapical stridulatory ridge. Protarsomere 5 without internal tooth. Modified protarsal claw with ventral tooth, not thickened. VARIATION. Male (n=59). Length 27.1-30.8 mm, width 16.8-17.9 mm (Fig. 61). Female (n=35). Length 29.5-31.9 mm, width 17.0-19.2
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Bulletin of the University of Nebra
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C O N T E N T S Abstract...........
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LIST OF FIGURES Fig. 1. Platycoelia
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Bulletin of the University of Nebra
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Members of the genus Platycoelia De
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alone to divide up the rutelines (a
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eing the most recent authority on t
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els Amongst the Great Andes of the
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tion. Zischka was a German who live
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Puncture size: large = 0.17 mm or l
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26. Pronotal surface sparsely punct
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Fig. 11. Platycoelia alternans male
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MONOGRAPHIC REVISION OF PLATYCOELIA
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Fig. 33. Platycoelia rufosignata me
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CNCI Canadian National Collection o
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subspecies AND varieties in the sam
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PHYLOGENETIC PATTERNS AND BIOGEOGRA
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metaunguitractor plate usually with
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larvae, and Paucar-Cabrera and Smit
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Erichson (1847) did not state how m
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species]; Ohaus 1918:178 [catalog l
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Platycoelia marginata and Platycoel
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tarsomere and metatarsomere 5 with
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5. Platycoelia meridensis new speci
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Page 66 and 67: MONOGRAPHIC REVISION OF PLATYCOELIA
Page 68 and 69: 7. Platycoelia prasina Erichson, 18
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Page 72 and 73: mentum. Apex of mentum with triangu
Page 74 and 75: 1972:302 [catalog listing as subspe
Page 76 and 77: green to yellow; parameres with the
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Page 86 and 87: VARIATION. Male (n=19). Length 19.2
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Page 96 and 97: gins. Ventrally with dark brown met
Page 98 and 99: 19. Platycoelia flavostriata (Latre
Page 102 and 103: 20. Platycoelia nigrosternalis Ohau
Page 104 and 105: LOCALITY DATA. 66 specimens examine
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Calloodes nitidissimus Lea, 1919 (C
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