Abai, MR

6th International Congress of DipterologyA phylogenetic study of the Bibionomorpha, with emphasisin the Mycetophiliformia (Diptera)Amorim, D.S. (1, 2) & E. Rindal (3)(1) FAPESP Grant n o 2003/10.274-9(2) Depto. de Biologia, FFCLRP, Universidade de São Paulo, Av. Bandeirantes 3900,14.040-901 Ribeirão Preto SP BRAZIL, dsamorim@usp.br(3) University of Oslo, Natural History Museum, Department of Zoology, P.O. Box 1172Blindern, 0318 Oslo, Norway, eirik.rindal@nhm.uio.noA phylogenetic analysis of the Mycetophiliformia (=Sciaroidea) wasperformed to establish relationships between families and to place generaof uncertain position in the system –Heterotricha, Ohakunea, Colonomyia,Freemanomyia, Rhynchoheterotricha, Chiletricha, Afrotricha, Anisotricha,Kenyatricha, Nepaletricha, Sciarosoma, Sciaropota, Insulatricha,Cabamofa, Rogambara, and Starkomyia. More than 150 species from 14families were examined and illustrated. A data matrix was built with 64terminals and 137 transformation series (202 character steps). A thoroughstudy of thorax morphology brought up robust data. Hennig’s system forthe Bibionomorpha was adopted, with Mycetophiliformia as a replacementname for Sciaroidea. The Anisopodiformia appeared as sister to the rest ofthe Bibionomorpha, followed by Pachyneuridae. Bibionidae appears assister to Mycetophiliformia. Within this clade Cecidomyiidae is sister tothe set of the remaining families, followed by the Sciaridae. The Sciaridaeappeared in all runs as a clade separated from the Mycetophilidae,Lygistorrhinidae or Rangomaramidae. The Rangomaramidae appeared asthe sister group of a clade consisting of (Ditomyiidae + Bolitophilidae +Diadocidiidae + Keroplatidae) and of (Lygistorrhinidae + Mycetophilidae).Rangomaramidae gathered most but not all genera of uncertain placement:a clade is constituted by Kenyatricha, Rhynchoheterotricha, Insularotricha,and Chiletricha; two separate clades include respectively Heterotricha andRangomarama; Ohakunea, Colonomyia, Cabamofa, and Rogambaraconstitute an additional stem in the family. Arachnocampa, usually placedin the Keroplatidae, does not appear within the keroplatids in most runs.Freemanomyia apparently fits as a basal clade in the Rangomaramidae.7