Abai, MR

6th International Congress of DipterologySystematic Study of the Genus Cladura Osten Sacken(Tipulidae) : Definitions and Diagnoses of its Species GroupsNakamura, T.Tochigi Prefectural Museum, Mutsumi-cho 2-2, Utsunomiya, Tochigi 320-0865, JapanThe genus Cladura Osten Sacken is a group of medium sized tipulidcraneflies, classified in the tribe Eriopterini of the subfamily Limoniinae,characterized by presence of cell m1 in wing venation. This genus issupposed to be one of the closest relatives of the flightless cranefly genusChionea Dalman. It is distributed in North America, Far East Russia, andEast Asia. Thirty-three species are described so far (five species in NorthAmerica, 26 in Japan and Russia, one in China, and one in Taiwan). Adultflies appear chiefly in late autumn.Recent study has revealed that the Japanese fauna of Cladura consists ofmore than 50 species, including at least 25 undescribed species. They canbe divided into five well distinguishable species groups, mentioned below.(1) C. decemnotata species group: Gonostylus with two apices.(2) C. nipponensis species group: Gonocoxite long and slender. Gonostylussimple, tapered apically, and hook shaped.(3) C. autumna species group: Ninth tergite with small tubercles at middleof posterior margin. Gonostylus with blunt tip.(4) C. megacauda species group: The largest group. Gonocoxite withapical prolongation and mesial projections. Gonostylus variable in shape.(5) C. machidella species group: Wing with supernumerary crossvein incells r3 and r4. Gonostylus broad hook shaped.C. japonica, the first described Japanese species, could not be assigned toany species groups, as it was described quite insufficiently with onlyfemale specimens.In this presentation, I will demonstrate the magnificent diversity of thegenus in Japan and the diagnoses of each species group, and discuss thephylogeny of these groups.Key Words: Cladura, Limoniinae, Tipulidae, species groups174