Abai, MR

6th International Congress of DipterologyFamily level phylogeny of the Carnoidea (Schizophora)Buck, M.Department of Environmental Biology, University of Guelph, Guelph, Ontario, Canada,N1G 2W1. E-mail: mbuck@uoguelph.ca.The Carnoidea is one of the most poorly defined superfamilies ofSchizophora. Its doubtful monophyly is reviewed and tentatively acceptedbased on one previously suggested and two newly discoveredsynapomorphies. The Carnoidea in its redefined sense includesAcartophthalmidae, Canacidae (including Tethinidae), Cryptochetidae,Milichiidae, Chloropidae, Carnidae, Australimyzidae and the recentlydescribed Inbiomyiidae. New evidence is provided for the paraphyly of'Tethinidae' with regard to Canacidae. The relationships of Carnoideafamilies are analyzed quantitatively for the first time based on a matrix of58 morphological characters, with Clusiidae and Heleomyzidae asoutgroups. The results of the analysis strongly support the previouslyhypothesized sister group relationship of Milichiidae and Chloropidae. Thelatter is not the sister group of Cryptochetidae as suggested by J.F.McAlpine (1989). Two other previously hypothesized sister grouprelationships, (1) Acarthophthalmidae + (Milichiidae + Chloropidae), and(2) Carnidae + Australimyzidae are not supported. Acartophthalmidae is abasal clade of Carnoidea but its sister group remains uncertain. Sistergroup of the Australimyzidae is the Inbiomyiidae. Separate family status ofAustralimyzidae (as opposed to subfamily status within Carnidae assuggested by some authors) is therefore necessary.Key Words: Carnoidea, phylogeny, Acartophthalmidae, Australimyzidae,Canacidae.∗∗∗∗∗∗∗∗∗∗33