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Lloyd Mycological Writings V4.pdf - MykoWeb

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THE STIPITATE STEREUMS.<br />

LEGEND. For some years we have been quite embarrassed by receiving many specimens of<br />

Stereums from foreign correspondents that we could not name, as we were familiar only with the<br />

species of Europe and the United States. During the past winter we studied by comparison, and by<br />

microscopical sections, the various historical specimens we found named in the several museums we<br />

visited (Kew, British Museum, Leiden, Berlin, and Paris). We found the Stereum species, as we find<br />

all mycological sections that we investigate, in a very chaotic and confused condition, owing to the<br />

multiplication of names and the careless and superficial work of those who have engaged in the promulgating<br />

of so-called "new species." Particularly is this true as regards the stipitate section of the<br />

genus, which, it appears to me, was in much worse condition (if possible) than the Apus section.<br />

In the earliest systematic work worthy of the name, Persoon's Synopsis Fungorum<br />

(1801), the Thelephoraceous plants, which are fungi with even hymenium, are<br />

divided into two genera, Thelephora and Merisma. The latter embraced the species<br />

of an encrusting nature, and while not so clearly defined is the same exactly as has<br />

in recent years been discovered to be a "new genus" and called "Soppittiella."<br />

The remainder of the plants which Persoon called Thelephora he divided into<br />

three sections<br />

Craterellus Pileus stipitate.<br />

Stereum -Pileus dimidiate.<br />

Corticium Resupinate.<br />

It will be noted that the plants embraced in this pamphlet would have been<br />

originally classed by Persoon under the sectional name Craterellus, and they may<br />

still be so juggled with as much merit as belongs to the most of such cheap work as<br />

is being done nowadays under the guise of "priority." In late years they have been<br />

called Podoscypha, also without much originality, as Persoon called the same section<br />

Craterellus.<br />

In the usually-employed Friesian system, which is only a modification of that<br />

of Persoon, Corticium was taken in nearly its original sense. Craterellus was restricted<br />

to the fleshy species and did not include either of the original species, and<br />

the remainder of the species were divided between Stereum and Thelephora.<br />

Exactly what distinction Fries had in mind between Stereum and Thelephora, it is<br />

hard to define. The "homogeneous" and "heterogeneous" nature of the tissue on<br />

which he based the difference is not marked enough to be the base for generic<br />

distinction. All of Fries' species of Stereum have hyaline spores and pale hymenium,<br />

and most of his species of Thelephora have colored spores and dark hymenium. It<br />

is the tendency of late years to make this the distinction between the two genera,<br />

and in my opinion it is the best. Many of the old species classed as Thelephora<br />

under this definition will fall into the genus Stereum.<br />

Modern "systematists" find it to their interest, of course, to break up the old<br />

genera into as many "new genera" as possible in order to make a lot of new names,<br />

which is the only advantage, and of very doubtful utility. Thus Karsten in 1881<br />

discovered that stipitate Stereums form a new genus, which he called Cotilydia.<br />

Patouillard in 1900 discovered that the same section wa? a "nov. gen.," which he<br />

called Podoscypha.<br />

There is another group of "systematists" who are engaged in discovering "new<br />

genera" on the " hairs " they find on the hymenium. Leveille was the father of these<br />

hair experts, but he was not particular as to the kind of "hairs." Any Thelephoraceous<br />

plant which had hairs was for Leveille a Hymenochaete, though he did not<br />

observe the subject closely enough to know which of his own species belonged to his<br />

"new genus." Cooke carried the matter further and divided the "hairs" into<br />

15

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