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Vines and Climbing Plants of Puerto Rico and the Virgin Islands

Vines and Climbing Plants of Puerto Rico and the Virgin Islands

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100<br />

<strong>Vines</strong> <strong>and</strong> <strong>Climbing</strong> <strong>Plants</strong> <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s<br />

1. Mikania congesta DC., Prodr. 5: 197. 1836.<br />

Fig. 35. A-D<br />

SYNONYMS: Mikania micrantha var. congesta (DC.) L. B.<br />

Rob.<br />

Mikania sc<strong>and</strong>ens sensu A. Stahl, non Willd.<br />

Guaco falso<br />

Herbaceous vine, twining, 2-5 m in length.<br />

Stems cylindrical or angular, striate, puberulous,<br />

or pilose. Leaves opposite; blades 2.5-14 × 1-10<br />

cm, deltate-ovate or oblong-ovate, chartaceous,<br />

with 3-5 main veins, <strong>the</strong> apex acute or acuminate,<br />

<strong>the</strong> base cordiform, <strong>the</strong> margins entire or weakly<br />

dentate or undulate; upper surface dull green,<br />

glabrous or puberulous; lower surface puberulous,<br />

densely covered with resinous dots; petioles 1.5-<br />

14 cm long, puberulous; pseudostipules forming<br />

a ring <strong>of</strong> lanceolate segments, early deciduous.<br />

Capitula numerous, sessile or short-pedunculate<br />

(4-5 mm long), arranged in dense, globose or<br />

corymbiform glomerules, terminal or at <strong>the</strong> end<br />

<strong>of</strong> short lateral branches; subinvolucral bract<br />

acicular, 2.5-4 mm long, puberulous; phyllaries<br />

oblong, 3-4 mm long, pilose, light green. Corollas<br />

white, infundibuliform, 2.5-3.5 mm long, with<br />

resinous dots at <strong>the</strong> apex. Achenes 3-3.5 mm long,<br />

black, cuneiform, with resinous dots; pappus <strong>of</strong><br />

33-37 bristles, white, scabrous, ca. 3 mm long.<br />

Phenology: Flowering <strong>and</strong> fruiting from<br />

November to March.<br />

Status: Native, relatively common.<br />

Distribution: Of wide distribution, in disturbed<br />

areas at lower <strong>and</strong> middle elevations. Also in<br />

Jamaica, Martinique, nor<strong>the</strong>rn South America,<br />

Peru, Bolivia, <strong>and</strong> Brazil.<br />

Public forests: Guánica, Maricao, Piñones,<br />

Río Abajo, <strong>and</strong> Susúa.<br />

Commentary: Mikania congesta is very<br />

similar to M. micrantha; never<strong>the</strong>less, it can be<br />

distinguished from M. micrantha by <strong>the</strong><br />

subinvolucral bracts usually <strong>of</strong> <strong>the</strong> same length<br />

or longer than <strong>the</strong> phyllaries (vs. shorter) <strong>and</strong> by<br />

<strong>the</strong> capitula arranged in subglobose glomerules<br />

(vs. capitula in corymbs).<br />

2. Mikania cordifolia (L.f.) Willd., Sp. Pl. 3:<br />

1746. 1804.<br />

Fig. 35. E-H<br />

BASIONYM: Cacalia cordifolia L.f.<br />

SYNONYMS: Mikania gonoclada DC.<br />

Mikania convolvulacea DC.<br />

Guaco<br />

Slightly woody vine, twining, 2-5 m in length.<br />

Stems pubescent, octagonal, striate. Leaves<br />

opposite; blades 7-12 × 3.5-7.4 (9) cm, ovate,<br />

slightly aromatic, chartaceous, <strong>the</strong> apex acute, <strong>the</strong><br />

base cordiform or subcordiform, <strong>the</strong> venation with<br />

three main veins, <strong>the</strong> margins serrate, upper<br />

surface dull green, puberulous; lower surface<br />

glaucous, densely pubescent, with prominent<br />

venation; petioles 5-4 cm long, sulcate;<br />

pseudostipules divided in lanceolate segments,<br />

whitish. Capitula numerous, arranged in axillary<br />

corymbiform cymes; subinvolucral bract linear to<br />

lanceolate, 4-5 mm long, pilose; phyllaries<br />

elliptical or narrowly ovate, 6-8 mm long, green,<br />

pubescent; corollas white, infundibuliform,<br />

aromatic; an<strong>the</strong>rs brown, connate, exserted.<br />

Achenes prismatic, with 5 longitudinal ribs, ca.<br />

3 mm long, grayish, scabrid; pappus <strong>of</strong> numerous<br />

white setae, scabrous, ca. 4.5 mm long.<br />

Phenology: Flowering <strong>and</strong> fruiting from<br />

November to March.<br />

Status: Native, very common.<br />

Distribution: In pastures or along rivers or<br />

roads, at middle <strong>and</strong> lower elevations. Also on<br />

Vieques <strong>and</strong> <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s; throughout <strong>the</strong><br />

Neotropics.<br />

Public forests: Carite, El Yunque, Guilarte,<br />

Maricao, Río Abajo, Toro Negro, Tortuguero, <strong>and</strong><br />

Vega.<br />

3. Mikania fragilis Urb., Symb. Antill. 1: 464.<br />

1899.<br />

Fig. 36. A-D<br />

SYNONYMS: Mikania fragilis Urb. var. leptodon Urb.<br />

Mikania rotunda sensu A. Stahl, non Griseb.<br />

Guaco redondo*<br />

Herbaceous vine, twining, 6-8 m in length.<br />

Stems angular, with numerous lateral branches,<br />

pubescent when young, grayish with numerous<br />

lenticels when mature, <strong>the</strong> inner bark reddish,<br />

aromatic. Leaves opposite; blade 5-11 × 4-10 cm,<br />

broadly ovate, fragile, fleshy, slightly aromatic,<br />

not gl<strong>and</strong>ular, <strong>the</strong> apex acute, obtuse, or rounded,<br />

<strong>the</strong> base cordiform, <strong>the</strong> margins serrate, revolute;<br />

upper surface dark green, shiny, with <strong>the</strong> reticulate<br />

venation sunken, scabrid; lower surface pale<br />

green, dull, sometimes with some resinous dots,<br />

venation prominent; petioles thick, 2-4 cm long,<br />

pubescent, sulcate; pseudostipules broadly ovate,

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