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Vines and Climbing Plants of Puerto Rico and the Virgin Islands

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<strong>Vines</strong> <strong>and</strong> <strong>Climbing</strong> <strong>Plants</strong> <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s 103<br />

reflexed, 8-10 mm wide. Capitula few,<br />

pedunculate, in axillary corymbiform cymes;<br />

bracts <strong>of</strong> <strong>the</strong> involucre lanceolate to narrowly<br />

ovate, ca. 12 mm long, green to whitish; corollas<br />

white or with a reddish or greenish tinge,<br />

infundibuliform; an<strong>the</strong>rs violet, connate, exserted.<br />

Achenes cuneiform, ca. 3.5 mm long, with 5<br />

longitudinal ribs, glabrous; pappus <strong>of</strong> numerous<br />

bristles, 7-8 mm long.<br />

Phenology: Flowering from September to April.<br />

Status: Endemic to <strong>Puerto</strong> <strong>Rico</strong>, ra<strong>the</strong>r common.<br />

Distribution: In forests <strong>of</strong> <strong>the</strong> cordilleras, at<br />

middle <strong>and</strong> upper elevations.<br />

Public forests: Carite, El Yunque, Guilarte,<br />

Maricao, <strong>and</strong> Toro Negro.<br />

* Common name taken from <strong>the</strong> work <strong>of</strong> Dr.<br />

Agustín Stahl (1937).<br />

4. Mikania micrantha Kunth in Humb. Bonpl.<br />

& Kunth, Nov. Gen. Sp. (quarto ed.) 4: 134.<br />

1818.<br />

Fig. 36. E-H<br />

Guaco falso<br />

Herbaceous vine, twining, attaining 10 m or<br />

more in length. Stems pubescent, angular, striate.<br />

Leaves opposite; blades 6-13 × 3-8 cm, ovate,<br />

chartaceous, 3-5-nerved, <strong>the</strong> apex acuminate, <strong>the</strong><br />

base cordiform, <strong>the</strong> margins dentate; upper surface<br />

slightly shiny, puberulous; lower surface<br />

puberulous, with <strong>the</strong> venation somewhat<br />

prominent; petioles sulcate, glabrous or<br />

puberulous, as long as or shorter than <strong>the</strong> blade;<br />

pseudostipules forming an interpetiolar ring <strong>of</strong><br />

lanceolate segments, early deciduous. Capitula<br />

numerous, pedunculate, arranged in corymbiform<br />

or paniculiform cymes, terminal on short lateral<br />

branches; subinvolucral bracts subulate, 2-3 mm<br />

long, glabrous; phyllaries ovate-oblong or<br />

elliptical, green, 3-4 mm long; corolla white,<br />

infundibuliform, slightly fragrant, 2.5-3 mm long;<br />

an<strong>the</strong>rs exserted. Achenes ca. 1.2 mm long, dark<br />

gray to black, pentagonal; pappus <strong>of</strong> numerous<br />

white bristles, 2-3 mm long.<br />

Phenology: Collected in flower <strong>and</strong> fruit from<br />

November to February.<br />

Status: Native, locally common.<br />

Distribution: In disturbed areas, such as<br />

pastures, along rivers, <strong>and</strong> marshy areas, at low<br />

elevation. Also throughout tropical America.<br />

Public forest: Maricao, Río Abajo, <strong>and</strong> Susúa.<br />

5. Mikania odoratissima Urb., Symb. Antill. 1:<br />

464. 1899.<br />

Fig. 37. A-C<br />

Slightly woody vine, twining, 3-5 m in length.<br />

Stems slender, striate, puberulous, aromatic.<br />

Leaves opposite; blades 4-10 × 3-8.5 cm,<br />

chartaceous, aromatic, ovate, triangular, or<br />

trilobed, glabrous or puberulous, 3-nerved, <strong>the</strong><br />

apex acute or acuminate, <strong>the</strong> base truncate to<br />

cordiform, <strong>the</strong> margins dentate; upper surface<br />

dark green, shiny, with <strong>the</strong> venation flat or slightly<br />

sunken; lower surface light green, dull, venation<br />

prominent; petioles 1.5-3.5 cm long, sulcate,<br />

puberulous; pseudostipules forming a ring <strong>of</strong><br />

lanceolate segments, pubescent, deciduous.<br />

Capitula in groups <strong>of</strong> 3, pedunculate to subsessile,<br />

arranged in terminal corymbiform cymes;<br />

subinvolucral bract obovate to oblong, 3-4 mm<br />

long; phyllaries 7-9 mm long, oblong-elliptical,<br />

green, puberulous; corollas white or pale green,<br />

infundibuliform, ca. 6.5 mm long. Achenes 4-4.5<br />

mm long, prismatic, scabrid, cream-colored;<br />

pappus <strong>of</strong> numerous scabrous bristles, creamcolored<br />

or light brown, 6.5-7 mm long.<br />

Phenology: Flowering <strong>and</strong> fruiting from<br />

November to March.<br />

Status: Endemic to <strong>Puerto</strong> <strong>Rico</strong>, uncommon.<br />

Distribution: In moist forests, at middle <strong>and</strong><br />

upper elevations, along <strong>the</strong> Cordillera Central.<br />

Public forests: Maricao <strong>and</strong> Río Abajo.<br />

6. Mikania pachyphylla Urb., Symb. Antill. 1:<br />

463. 1899.<br />

Fig. 37. D-G<br />

Slightly woody vine, twining, 4-8 m in length.<br />

Stems hexagonal, 8-10 mm in diameter, glabrous,<br />

with numerous lateral branches, <strong>the</strong> bark grayish,<br />

smooth, <strong>and</strong> aromatic. Leaves opposite; blades<br />

3.5-9 × 2-3 cm, ovate, fleshy, brittle, slightly<br />

aromatic, 3-nerved, <strong>the</strong> apex acuminate, <strong>the</strong> base<br />

obtuse or rounded, <strong>the</strong> margins serrate-mucronate;<br />

upper surface pale green, shiny, glabrous, foveate,<br />

with <strong>the</strong> venation sunken; lower surface pale<br />

green, dull, glabrous, with prominent venation<br />

<strong>and</strong> whitish; petioles 1-2 cm long, glabrous,

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