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Vines and Climbing Plants of Puerto Rico and the Virgin Islands

Vines and Climbing Plants of Puerto Rico and the Virgin Islands

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22<br />

<strong>Vines</strong> <strong>and</strong> <strong>Climbing</strong> <strong>Plants</strong> <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s<br />

1. Family BLECHNACEAE<br />

1. BLECHNUM<br />

Terrestrial ferns, hemiepiphytic or less frequently epiphytic; rhizomes elongate, erect or sc<strong>and</strong>ent,<br />

sometimes forming a caudex or stolon, densely covered with scales. Fronds sterile or fertile, monomorphic<br />

or dimorphic; blades glabrous, pinnatisect, simple-pinnate or less frequently entire; veins free, bifurcated,<br />

except for <strong>the</strong> minor veins on <strong>the</strong> fertile fronds; pinnae entire or serrate. Sori linear, parallel to <strong>the</strong><br />

midvein; indusia linear, opening toward <strong>the</strong> midvein. A cosmopolitan genus <strong>of</strong> about 180 species,<br />

mostly in <strong>the</strong> Sou<strong>the</strong>rn Hemisphere.<br />

1. Blechnum fragile (Liebm.) C. V. Morton &<br />

Lellinger, Amer. Fern J. 57: 68. 1967.<br />

Fig. 5. A-C<br />

BASIONYM: Lomaria fragilis Liebm.<br />

Slightly woody fern, hemiepiphytic, sc<strong>and</strong>ent<br />

by means <strong>of</strong> adventitious roots, attaining 2 m in<br />

length. Rhizomes thick, 0.8-1.5 cm wide, densely<br />

covered with subulate, slihtly undulate, ciliate,<br />

ferruginous scales, 8-17 mm long. Fronds more<br />

or less clustered, arching, ascendent or pendulous,<br />

dimorphic; blades pinnate, membranaceous.<br />

Sterile fronds 40-60 × 4-12 cm, with an elliptical<br />

outline, <strong>the</strong> apex acuminate or caudate, <strong>the</strong> base<br />

acute (<strong>the</strong> pinnae gradually reduced in size toward<br />

<strong>the</strong> base); pinnae 25-75 pairs, oblong-deltate, <strong>the</strong><br />

apex acuminate, <strong>the</strong> margins revolute, sometimes<br />

with minute callus-like projections; <strong>the</strong> upper<br />

2. Family DAVALLIACEAE<br />

1. OLEANDRA<br />

surface dull, with a prominent midvein; <strong>the</strong> lower<br />

surface dull, paler than <strong>the</strong> upper surface, with a<br />

prominent midvein; stipe 10-18 cm long, blackish<br />

or brown, with some vestigial pinnae. Fertile<br />

fronds slightly smaller than <strong>the</strong> sterile ones;<br />

pinnae 28-41 pairs, linear; indusia half as wide<br />

as <strong>the</strong> pinna.<br />

Phenology: Found in fertile condition<br />

throughout <strong>the</strong> year.<br />

Status: Native, common.<br />

Distribution: In moist forests at middle <strong>and</strong><br />

upper elevations along <strong>the</strong> Cordillera Central <strong>and</strong><br />

<strong>the</strong> Sierra de Luquillo. Also in Jamaica, Cuba,<br />

Hispaniola, sou<strong>the</strong>rn Mexico, <strong>and</strong> Central<br />

America.<br />

Public Forests: Carite, El Yunque, Guilarte,<br />

<strong>and</strong> Toro Negro.<br />

Epiphytic or terrestrial ferns with very elongate rhizomes, erect, creeping, or climbing, sometimes<br />

branched, densely covered with peltate scales. Fronds sterile or fertile, monomorphic; blades simple,<br />

entire; veins free <strong>and</strong> parallel; stipe articulate. Sori circular, usually adjacent to <strong>the</strong> midvein; indusia<br />

circular or reniform, peltate. A tropical genus <strong>of</strong> about 40 species.<br />

1. Ole<strong>and</strong>ra articulata (Sw.) C. Presl, Tent.<br />

Pterid. 78. 1836.<br />

Fig. 5. D-E<br />

BASIONYM: Aspidium articulatum Sw.<br />

SYNONYM: Ole<strong>and</strong>ra nodosa (Willd.) C. Presl<br />

Herbaceous fern, epiphytic or climbing by<br />

adventitious roots, attaining 5 m in length.<br />

Rhizomes slender, densely covered with<br />

lanceolate, ferruginous scales. Fronds pendulous;<br />

blades (15) 23-39 (42) × (2.5) 3-6 cm,<br />

membranaceous, oblong, elliptical, or<br />

sublanceolate, <strong>the</strong> apex acute, acuminate, or<br />

caudate, <strong>the</strong> base acute or acuminate, sometimes<br />

unequal, <strong>the</strong> margin crenulate or crenate, slightly<br />

revolute <strong>and</strong> ferruginous; <strong>the</strong> upper surface shiny,

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