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Vines and Climbing Plants of Puerto Rico and the Virgin Islands

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<strong>Vines</strong> <strong>and</strong> <strong>Climbing</strong> <strong>Plants</strong> <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s 321<br />

1. Bougainvillea glabra Choisy in DC., Prodr.<br />

13(2): 437. 1849.<br />

Fig. 129. A-F<br />

Trinitaria<br />

Woody vine or shrub, clambering, attainig 15<br />

m in length. Stem cylindrical, attaining 5-6 cm<br />

in diameter; cross section <strong>of</strong> <strong>the</strong> stem with discrete<br />

vascular bundles within <strong>the</strong> connective tissue.<br />

Branches pubescent, puberulent, or glabrous, with<br />

straight, axillary spines. Leaves alternate or<br />

subopposite, 3-6 × 1.5-3 cm, chartaceous,<br />

elliptical, puberulent, <strong>the</strong> apex acuminate, <strong>the</strong> base<br />

attenuate, slightly asymmetrical, <strong>the</strong> margins<br />

slightly sinuate; lower surface with <strong>the</strong> midvein<br />

prominent; petioles slender, 4-12 mm long.<br />

Dichasial cymes compound, on axillary branches<br />

or at <strong>the</strong> end <strong>of</strong> axillary spines, <strong>the</strong> axes glabrous<br />

or puberulent; bracts ovate, glabrous, cordiform<br />

at <strong>the</strong> base, 2.5-4 cm long, <strong>of</strong> showy colors, red,<br />

deep pink, orange, white, or purple. Perianth<br />

tubular-hypocrateriform, 2.5-3 mm long,<br />

pubescent, with 8-10 obtuse lobes, cream-colored.<br />

Anthocarp fusiform, glabrous, ca. 1.5 cm long.<br />

Phenology: Flowering throughout <strong>the</strong> year.<br />

Status: Exotic, very common in our gardens.<br />

Distribution: Much cultivated throughout<br />

<strong>Puerto</strong> <strong>Rico</strong>. This is <strong>the</strong> most common species <strong>of</strong><br />

its genus in <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s. It<br />

grows well in dry areas <strong>and</strong> requires little<br />

watering, for which reasons it is cultivated along<br />

3. PISONIA<br />

avenues <strong>and</strong> highways. Also throughout <strong>the</strong><br />

Antilles. Native to South America, but widely<br />

cultivated around <strong>the</strong> globe.<br />

2. Bougainvillea spectabilis Willd., Sp. Pl. ed.<br />

2, 348. 1799.<br />

Fig. 129. G-H<br />

Trinitaria<br />

Woody shrub, erect or clambering, attainig 5<br />

m in length. Branches pilose, with straight,<br />

axillary, pilose spines. Leaves alternate, 2.6-2.5<br />

× 2.7-5 cm, chartaceous, ovate, <strong>the</strong> apex acute,<br />

obtuse, or acuminate, <strong>the</strong> base obtuse, rounded,<br />

or attenuate, slightly asymmetrical, <strong>the</strong> margins<br />

sinuate; lower surface tomentulose, with<br />

prominent pilose venation; petioles slender, 1-2.3<br />

cm long, pilose. Dichasial cymes compound, on<br />

axillary branches or at <strong>the</strong> end <strong>of</strong> axillary spines,<br />

<strong>the</strong> axes pilose; bracts ovate, pubescent, pilose<br />

along <strong>the</strong> midvein, cordiform at <strong>the</strong> base, 3.6.5.5<br />

cm long, red or deep pink. Perianth tubularhypocrateriform,<br />

2.2-3 mm long, pilose.<br />

Phenology: Flowering throughout <strong>the</strong> year.<br />

Status: Exotic, uncommon.<br />

Distribution: Little cultivated in <strong>Puerto</strong> <strong>Rico</strong>.<br />

Also throughout <strong>the</strong> Antilles. Native to South<br />

America, but widely cultivated on a worldwide<br />

level.<br />

Trees, shrubs, or lianas, dioecious or monoecious. Leaves opposite or subopposite, sometimes with<br />

axillary spines. Flowers unisexual, small, white or pale yellow, in axillary or terminal panicles, corymbs,<br />

or cymes; bracts <strong>and</strong> bracteoles minute, persistent. Staminate flowers campanulate to obconical; stamens<br />

6-8, exserted, <strong>the</strong> filaments unequal, connate at <strong>the</strong> base. Pistillate flowers tubular, 5-lobate; ovary<br />

superior, sessile, <strong>the</strong> style exserted, <strong>the</strong> stigma branched. Fruit a dry anthocarp, oblanceolate or ellipsoid,<br />

with 5 ribs that bear sticky, stipitate gl<strong>and</strong>s. A genus <strong>of</strong> 10-15 species, <strong>of</strong> tropical distribution.<br />

1. Pisonia aculeata L., Sp. Pl. 1026. 1753.<br />

Fig. 1A; 128. G-M<br />

SYNONYM: Pisonia helleri St<strong>and</strong>l.<br />

Escambrón, Prickly-mampoo<br />

Liana or woody, clambering shrub, attainig<br />

20 m in length. Stems more or less cylindrical,<br />

glabrous or minutely pubescent, grayish or cream-<br />

colored, lenticellate, attaining 7 cm in diameter;<br />

cross section <strong>of</strong> <strong>the</strong> stem with discrete vascular<br />

bundles within <strong>the</strong> connective tissue. Branches<br />

short or elongate; short branches opposite,<br />

divaricate, persistent, with recurved spines,<br />

produced on <strong>the</strong> trunk; elongate branches at <strong>the</strong><br />

end <strong>of</strong> <strong>the</strong> stem, glabrescent, quadrangular, with<br />

recurved axillary spines. Leaves opposite on <strong>the</strong><br />

elongate branches or whorled on <strong>the</strong> short lateral

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