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Vines and Climbing Plants of Puerto Rico and the Virgin Islands

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<strong>Vines</strong> <strong>and</strong> <strong>Climbing</strong> <strong>Plants</strong> <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s 285<br />

7a. Stipules truncate at <strong>the</strong> base; peduncle <strong>of</strong> <strong>the</strong> inflorescence as long as or shorter than <strong>the</strong> accompanying<br />

leaf; legume oblong, 8-9 mm wide, curved in <strong>the</strong> middle...................…………..1. V. adenantha<br />

7b. Stipules asymmetrical, <strong>the</strong> base auriculate on only one side; legume linear, curved in <strong>the</strong> distal<br />

portion, 3-4 mm wide…….............…….…………………………………..….2. V. antillana<br />

8a. Plant more or less glabrous; legume 10-40 cm long …………………..….8. V. unguiculata<br />

8b. Plant ferruginous-pilose; legume 7-11 cm long ………………………….......9. V. vexillata<br />

1. Vigna adenantha (G. Meyer) Marechal,<br />

Mascherpe & Stanier, Taxon 27: 202. 1978.<br />

Fig.114. E-H<br />

BASIONYM: Phaseolus adenanthus G. Meyer<br />

Habichuela cimarrona<br />

Slightly woody vine, twining, climbing,<br />

attainig 3-5 m in length. Stems slender, almost<br />

cylindrical, glabrous or pilose. Leaves alternate,<br />

trifoliolate; leaflets chartaceous, with three<br />

principal veins from <strong>the</strong> base, <strong>the</strong> apex acute,<br />

obtuse, or acuminate, <strong>the</strong> upper surface glabrous,<br />

sometimes sparsely appressed-pubescent, <strong>the</strong><br />

lower surface pale green, appressed-pubescent,<br />

especially on <strong>the</strong> prominent veins, <strong>the</strong> margins<br />

deeply sinuate; terminal leaflet lanceolate or ovate,<br />

3-9.5(15.5) × 1.5-5.3(9.6) cm, <strong>the</strong> base cuneate<br />

or obtuse; lateral leaflets markedly asymmetrical,<br />

with one side elliptical <strong>and</strong> <strong>the</strong> o<strong>the</strong>r ovate or<br />

lanceolate, <strong>the</strong> base cuneate-obtuse; petiolules<br />

pubescent, 2-5 mm long; rachis 1-2 cm long,<br />

striate, pilose; petioles canaliculate, striate, pilose<br />

or glabrous, 3-9.5 cm long; stipules triangular,<br />

3-4 mm long, glabrous, conspicuously veined,<br />

truncate at <strong>the</strong> base; stipels oblong, ca. 2 mm long.<br />

Inflorescence <strong>of</strong> axillary pseudoracemes, 4-12 cm<br />

long, with 2 flowers per node; peduncle appressedpubescent<br />

or glabrous; pedicels 1.6-2 mm long,<br />

glabrous; bracts in pairs, lanceolate, deciduous.<br />

Calyx green, glabrous, campanulate, 5-7.5 mm<br />

long, with 3 lanceolate lobes, longer than <strong>the</strong> two<br />

central ones, which are rounded <strong>and</strong> fused; corolla<br />

pale violet; st<strong>and</strong>ard reniform, retuse at <strong>the</strong> apex,<br />

2.2-3 cm long <strong>and</strong> up to 2.5 cm wide, pale violet,<br />

whitish toward <strong>the</strong> margins, with <strong>the</strong> vexillar<br />

callus yellow; wings 2.2-2.5 cm long, obovate,<br />

twisted, enclosing <strong>the</strong> keel, pale violet, dark violet<br />

toward <strong>the</strong> apex, or sometimes pale yellow; keel<br />

up to 3 cm wide, pale violet to whitish, spirally<br />

twisted at <strong>the</strong> apex (½-1½ turns). Legume oblong,<br />

curved, flattened, <strong>the</strong> area <strong>of</strong> <strong>the</strong> seeds slightly<br />

prominent, 7-11 cm long, 8-9 mm wide,<br />

puberulent, scabrous along <strong>the</strong> margins, <strong>the</strong> apex<br />

acute, recurved. Seeds 11-16, almost circular,<br />

flattened, ca. 6 mm wide, dark brown.<br />

Phenology: Collected in flower <strong>and</strong> fruit from<br />

December to April.<br />

Status: Probably native, ra<strong>the</strong>r common.<br />

Distribution: Moist disturbed areas at lower<br />

<strong>and</strong> middle elevations. Native to tropical America,<br />

cultivated in Africa <strong>and</strong> Asia.<br />

Public Forests: El Yunque, Guajataca, <strong>and</strong> Río<br />

Abajo.<br />

2. Vigna antillana (Urb.) Fawcett & Rendle, Fl.<br />

Jam. 4(2): 69. 1920.<br />

Fig. 114. A-D<br />

BASIONYM: Phaseolus antillanus Urb.<br />

Herbaceous vine, twining, climbing, attainig<br />

3-5 m in length. Stems cylindrical, glabrous,<br />

reddish on <strong>the</strong> young portions. Leaves alternate,<br />

trifoliolate; leaflets chartaceous, 3-9 × 2-7 cm,<br />

<strong>the</strong> apex acuminate, <strong>the</strong> margins entire, slightly<br />

revolute; upper surface glabrous or puberulent,<br />

with slightly prominent venation; lower surface<br />

glabrous or puberulent, with prominent venation;<br />

terminal leaflet broadly ovate or deltate, <strong>the</strong> base<br />

cuneate or truncate, with three main veins; lateral<br />

leaflets markedly asymmetrical, with one side<br />

oblong <strong>and</strong> <strong>the</strong> o<strong>the</strong>r broadly ovate, <strong>the</strong> base<br />

truncate; petiolules puberulent, 4-5 mm long;<br />

rachis 2-2.5 cm long, marginate, puberulent or<br />

glabrous; petioles marginate, puberulent or<br />

glabrous, 3-10 cm long; stipules lanceolate, 6-8<br />

mm long, asymmetrical at <strong>the</strong> base, one side<br />

auriculate; stipels subulate, minute, persistent.<br />

Inflorescence <strong>of</strong> axillary pseudoracemes, up to 30<br />

cm long, with 2 flowers per node; peduncle<br />

glabrous or puberulent; pedicels 2-3 mm long,<br />

glabrous; bracts minute, persistent. Calyx green,<br />

puberulent, campanulate, 4-5 mm long, <strong>the</strong> lobes<br />

more or less <strong>of</strong> <strong>the</strong> same length, 3 <strong>of</strong> <strong>the</strong>m<br />

triangular, <strong>the</strong> 2 central ones rounded <strong>and</strong> fused;

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