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Vines and Climbing Plants of Puerto Rico and the Virgin Islands

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258<br />

<strong>Vines</strong> <strong>and</strong> <strong>Climbing</strong> <strong>Plants</strong> <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s<br />

<strong>the</strong> base obtuse or rounded; petiolules 2-5 mm<br />

long, pilose; petioles 4.5-10.5 cm long, uncinatepubescent,<br />

striate; rachis 1-6 mm long, pilose,<br />

canaliculate; stipules ovate to lanceolate, 5-10 mm<br />

long, deciduous; stipels filiform, ca. 2 mm long.<br />

Inflorescences <strong>of</strong> axillary racemes, erect, 30-45<br />

cm long, <strong>the</strong> axis pubescent, dark brown, with<br />

flowers paired along its length; pedicels 12-17<br />

mm long. Calyx purple, campanulate, ca. 2 mm<br />

long, pilose; corolla pink or pale violet, ca. 4 mm<br />

long. Legume uncinate-pubescent, 0.7-1(2) cm<br />

long, crenate on <strong>the</strong> ventral suture, with (1)2(3)<br />

fertile segments. Seeds ca. 3.5 mm long, oblongelliptical,<br />

light brown.<br />

Phenology: Collected in flower <strong>and</strong> fruit from<br />

January to August.<br />

Status: Native, ra<strong>the</strong>r common.<br />

Distribution: On <strong>the</strong> ground in dry to moist<br />

forests <strong>and</strong> in areas <strong>of</strong> disturbed vegetation along<br />

<strong>the</strong> Cordillera Central. Also on St. Croix;<br />

distributed throughout <strong>the</strong> Antilles, Central <strong>and</strong><br />

South America.<br />

Public Forests: El Yunque, Guilarte, Maricao,<br />

Río Abajo, <strong>and</strong> Toro Negro.<br />

Key to <strong>the</strong> varieties in <strong>Puerto</strong> <strong>Rico</strong><br />

1a. Leaflets acute or acuminate at <strong>the</strong> apex....... 2<br />

1b. Leaflets obtuse… D. axillare var. axillare<br />

2a. <strong>Plants</strong> densely pubescent ........……………….<br />

.…………… D. axillare var. acutifolium<br />

2b. <strong>Plants</strong> puberulent..........……………………<br />

........…………. D. axillare var. stoloniferum<br />

2. Desmodium incanum DC., Prodr. 2: 332.<br />

1825.<br />

Fig. 103. D-G<br />

SYNONYM: Hedysarum canum J.F. Gmel.<br />

Desmodium canum (J.F. Gmel.) Schinz & Thell.<br />

Hedysarum supinum Sw.<br />

Meibomia supina (Sw.) Britton<br />

Decumbent or sc<strong>and</strong>ent herb, much branched<br />

from <strong>the</strong> woody base, attainig 1-1.5 m in length.<br />

Stems more or less cylindrical, appressedpubescent<br />

or pilose, glabrescent with age. Leaves<br />

alternate, trifoliolate; leaflets 3-9 × 0.8-3.8 cm,<br />

elliptical, oblong, ovate, lanceolate, or<br />

exceptionally rounded, chartaceous, involute, <strong>the</strong><br />

apex obtuse or acute, mucronate, <strong>the</strong> base obtuse<br />

or rounded, <strong>the</strong> margins entire; upper surface dark<br />

green, puberulent; lower surface grayish green,<br />

strigulose, with prominent venation; petiolules<br />

1.5-3 mm long, pilose; petioles 0.7-3 cm long,<br />

pilose, canaliculate; rachis 3-10 mm long, pilose,<br />

canaliculate; stipules ovate to lanceolate, 6-10 mm<br />

long, persistent; stipels minute, subulate.<br />

Inflorescences <strong>of</strong> terminal pseudoracemes, 10-18<br />

cm long, pubescent; bracts minute, persistent.<br />

Calyx green, campanulate, 2.5-3 mm long,<br />

strigulose; corolla pink or pink-violet, <strong>the</strong><br />

st<strong>and</strong>ard ovate, 5-6 mm long, retuse, unguiculate<br />

at <strong>the</strong> base, <strong>the</strong> wings <strong>and</strong> <strong>the</strong> keel as long as <strong>the</strong><br />

st<strong>and</strong>ard. Legume uncinate-pubescent, 2-4 cm<br />

long, crenate on <strong>the</strong> ventral suture, with 5-8 fertile<br />

segments. Seeds 2.5-3.5 mm long, oblongelliptical,<br />

light brown.<br />

Phenology: Flowering <strong>and</strong> fruiting throughout<br />

<strong>the</strong> year.<br />

Status: Apparently native, very common.<br />

Distribution: In <strong>the</strong> understory <strong>of</strong> dry to moist<br />

forests <strong>and</strong> in areas <strong>of</strong> disturbed vegetation. Also<br />

on Mona, Vieques, St. Croix, St. John, St. Thomas,<br />

Tortola, <strong>and</strong> <strong>Virgin</strong> Gorda; distributed throughout<br />

<strong>the</strong> tropics as a weed.<br />

Public Forests: El Yunque, Guajataca,<br />

Guánica, Maricao, Mona, Río Abajo, <strong>and</strong> Susúa.<br />

3. Desmodium intortum (Mill.) Urb., Symb.<br />

Antill. 8: 292. 1920.<br />

Fig. 103. H-M<br />

BASIONYM: Hedysarum intortum Mill.<br />

Decumbent or ascendent herb, branched from<br />

<strong>the</strong> base, attainig 3 m in length. Stems trigonal,<br />

densely covered with uncinate hairs. Leaves<br />

alternate, trifoliolate; leaflets 4-7 × 2-5 cm, ovate,<br />

broadly ovate, or elliptical, chartaceous, <strong>the</strong> apex<br />

obtuse or acute, <strong>the</strong> base rounded, <strong>the</strong> margins<br />

entire; upper surface dark green, strigulose; lower<br />

surface pale green, more or less strigose, with<br />

prominent venation; petiolules 2.5-3 mm long,<br />

densely pilose; petioles 2-3.5 cm long, pilose,<br />

striate; rachis ca. 5 mm long, pilose; stipules<br />

ovate, acuminate, ca. 7 mm long, deciduous;<br />

stipels minute, subulate. Inflorescences <strong>of</strong><br />

terminal or axillary pseudoracemes, uncinatepubescent.<br />

Calyx green, campanulate, puberulent,<br />

<strong>the</strong> tubular portion ca. 3 mm long, <strong>the</strong> inner sepal<br />

ca. 3.5 mm long, lanceolate, <strong>the</strong> rest ca. 1.5 mm<br />

long, deltate; corolla pink-violet, <strong>the</strong> st<strong>and</strong>ard ca.

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