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Vines and Climbing Plants of Puerto Rico and the Virgin Islands

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390<br />

<strong>Vines</strong> <strong>and</strong> <strong>Climbing</strong> <strong>Plants</strong> <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s<br />

1. Clerodendrum aculeatum (L.) Schltdtl.,<br />

Linnaea 6: 750. 1831.<br />

Fig. 159. A-E<br />

BASIONYM: Volkameria aculeata L.<br />

Erect or sc<strong>and</strong>ent shrub attainig 3 m in length.<br />

Stems cylindrical, usually striate, puberulent, with<br />

2 or 3 spines, opposite or whorled, in <strong>the</strong> area <strong>of</strong><br />

<strong>the</strong> nodes; stipules absent. Leaves opposite or three<br />

per node, 1.5-7 × 0.5-2.6 cm, oblong or ellipticovate,<br />

chartaceous, glabrous or puberulent, <strong>the</strong><br />

apex acute or obtuse, <strong>the</strong> base cuneate or rounded,<br />

<strong>the</strong> margins entire; upper surface dull, dark green;<br />

lower surface pale green, dull, with numerous<br />

49. Family VERBENACEAE<br />

Key to <strong>the</strong> genera<br />

1a. Flowers sessile, subtended by an involucre <strong>of</strong> 3(4) elongate bracts, exp<strong>and</strong>ed, pink or lilac at <strong>the</strong><br />

base, much longer than <strong>the</strong> calyx.………………………………..….……..2. Congea<br />

1b. Flowers with minute bracts, not covering <strong>the</strong> calyx……………………...………………..2<br />

2.a Calyx gamosepalous, acetabuliform, orange, <strong>the</strong> sepals not apparent ….….3. Holmskioldia<br />

2b. Calyx <strong>of</strong> free sepals or sepals connate only at <strong>the</strong> base, <strong>of</strong> various colors, but not orange…………....3<br />

3a. Inflorescences corymbiform; calyx green, white, or pink, <strong>the</strong> sepals imbricate……1. Clerodendrum<br />

3b. Inflorescences racemose; calyx violet, <strong>the</strong> sepals exp<strong>and</strong>ed .................……………….…....4. Petrea<br />

1. CLERODENDRUM<br />

Trees, erect shrubs, or less frequently lianas or sc<strong>and</strong>ent shrubs, sometimes with axillary spines.<br />

Leaves simple, opposite or whorled, petiolate, with entire or dentate margins. Flowers showy, bisexual,<br />

zygomorphic, in axillary cymes or terminal panicles. Calyx campanulate or tubular, truncate, 5-lobate;<br />

corolla 5-merous, hypocrateriform; stamens 4, <strong>the</strong> filaments didynamous, adnate to <strong>the</strong> base <strong>of</strong> <strong>the</strong><br />

corolla, exserted; ovary superior, 4-locular, each locule with a solitary ovule; style slender, crowned by<br />

a globose stigma. Fruit drupaceous, 4-lobate, with 4 pyrenes. A genus <strong>of</strong> approximately 400 species,<br />

<strong>the</strong> majority native to <strong>the</strong> tropics <strong>of</strong> <strong>the</strong> Old World.<br />

Key to <strong>the</strong> species <strong>of</strong> Clerodendrum<br />

1a. Branches with conical spines, opposite or whorled, in <strong>the</strong> area <strong>of</strong> <strong>the</strong> nodes................1. C. aculeatum<br />

1b. Branches unarmed....…………………………………………………………………………2<br />

2a. Calyx reddish, 1-1.5 cm long …............……….………..……...……………...….. C. × speciosum<br />

2b. Calyx white, 1.5-2.5 cm long ........….........………………..………………..…………. C. thomsonae<br />

dots; petioles 2.5-10 mm long, puberulent.<br />

Inflorescences <strong>of</strong> axillary cymes. Calyx<br />

campanulate, puberulent, 2.5-4 mm long, <strong>the</strong><br />

sepals exp<strong>and</strong>ed; corolla white, hypocrateriform,<br />

1.4-2.7 cm long, <strong>the</strong> tube sometimes with a purple<br />

tinge; filaments pink, twice as long as <strong>the</strong> corolla;<br />

style purple, as long as <strong>the</strong> filaments. Drupe ovoid<br />

or depressed-ovoid, 5-7 mm long, opening in two<br />

halves when ripe.<br />

Phenology: Collected in flower from January<br />

to March <strong>and</strong> from August to October <strong>and</strong> in fruit<br />

from January to August.<br />

Status: Native, locally common.

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