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Vines and Climbing Plants of Puerto Rico and the Virgin Islands

Vines and Climbing Plants of Puerto Rico and the Virgin Islands

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<strong>Vines</strong> <strong>and</strong> <strong>Climbing</strong> <strong>Plants</strong> <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s<br />

Public Forests: Carite, El Yunque, Guánica,<br />

Guajataca, Maricao, Río Abajo, Susúa, Toro<br />

Negro, <strong>and</strong> Tortuguero.<br />

Commentary: In <strong>the</strong> first edition <strong>of</strong> this work,<br />

Smilax coriacea was considered a synonym <strong>of</strong> S.<br />

havanensis Jacq. This error is rectified in this<br />

edition, since <strong>the</strong> two species are different <strong>and</strong><br />

can be distinguished by <strong>the</strong>ir pattern <strong>of</strong> <strong>the</strong><br />

venation. The secondary veins on <strong>the</strong> leaves <strong>of</strong> S.<br />

coriacea form an angle with <strong>the</strong> principal vein<br />

that varies from 45º to 90º, while in S. havanensis<br />

<strong>the</strong> secondary veins form an angle with <strong>the</strong><br />

principal vein that varies from 25º to 35º. Smilax<br />

havanensis is found in Cuba <strong>and</strong> in Hispaniola,<br />

but it is not found in <strong>Puerto</strong> <strong>Rico</strong>, nor in <strong>the</strong> <strong>Virgin</strong><br />

Isl<strong>and</strong>s.<br />

2. Smilax domingensis Willd, Sp. Pl. 4: 783.<br />

1806.<br />

Fig. 184. G-K<br />

Bejuco de membrillo, Raíz de zarzaparilla,<br />

Zarzaparilla<br />

Slightly woody vine that climbs by means <strong>of</strong><br />

tendrils <strong>and</strong> attains 5-10 m in length. Stems<br />

slender, strong, glabrous, smooth, scarcely<br />

branched. Leaves alternate, simple, chartaceous,<br />

5-12(15) × 2-6.5 cm, ovate, oblong, elliptical, or<br />

broadly elliptical, usually involute, 5-veined, <strong>the</strong><br />

apex acuminate, <strong>the</strong> base obtuse, rounded, or less<br />

frequently subcordate; <strong>the</strong> margins entire; upper<br />

surface shiny, glabrous; lower surface dull,<br />

glabrous, papillose, with prominent venation;<br />

petioles 1-1.5 cm long, articulated at <strong>the</strong> apex,<br />

forming a sheath at <strong>the</strong> base, from which two<br />

simple tendrils are borne, 8-12 cm long.<br />

Inflorescence axillary, umbelliform, solitary.<br />

Flowers greenish, fragrant. Staminate flowers<br />

with pedicels 5-9 mm long; tepals elliptical, 3-<br />

3.5 mm long. Fruit globose, fleshy, ca. 1 cm in<br />

diameter, dark violet or almost black, with a single<br />

seed inside.<br />

Phenology: Flowering from May to August<br />

<strong>and</strong> fruiting from December to February.<br />

Status: Native, ra<strong>the</strong>r common.<br />

Distribution: In moist forests at middle to<br />

upper elevations, along <strong>the</strong> Cordillera Central <strong>and</strong><br />

in <strong>the</strong> zone <strong>of</strong> mogotes. Also in <strong>the</strong> Greater<br />

Antilles, Mexico, Guatemala, <strong>and</strong> Belize.<br />

Public Forests: Cambalache, El Yunque,<br />

Guajataca, Maricao, Río Abajo, Susúa, <strong>and</strong><br />

Tortuguero.<br />

Reference: Acevedo-Rodríguez, P. (in prep.) Smilacaceae. In: Acevedo-Rodríguez, P. & M. T. Strong<br />

(eds.) Monocots <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s. Contrib. U.S. Natl. Herb.

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