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Vines and Climbing Plants of Puerto Rico and the Virgin Islands

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122<br />

<strong>Vines</strong> <strong>and</strong> <strong>Climbing</strong> <strong>Plants</strong> <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s<br />

Phenology: Flowering <strong>and</strong> fruiting almost<br />

throughout <strong>the</strong> year, especially from November<br />

to July.<br />

Status: Native, very common in southwestern<br />

<strong>Puerto</strong> <strong>Rico</strong>.<br />

5. MACFADYENA<br />

Lianas with tendrils. Stems cylindrical or quadrangular; cross section <strong>of</strong> <strong>the</strong> mature stem with<br />

multilobed xylem, <strong>the</strong> lobes alternating with radially arranged phloem tissue; gl<strong>and</strong>ular interpetiolar<br />

zone present or absent; pseudostipules small, lanceolate to ovate. Leaves opposite, 2-foliolate, with a<br />

terminal trifid tendril in <strong>the</strong> form <strong>of</strong> a claw. Flowers in axillary cymes or panicles. Calyx campanulate,<br />

simple; corolla yellow, zygomorphic, tubular or campanulate; stamens 4, didynamous; ovary superior,<br />

linear-cylindrical, with 2 locules <strong>and</strong> an annular disc at <strong>the</strong> base. Fruit a narrow capsule, elongate <strong>and</strong><br />

compressed, with <strong>the</strong> valves parallel to <strong>the</strong> septum; seeds numerous, with 2 wings. A genus <strong>of</strong> 4 species<br />

<strong>of</strong> neotropical distribution.<br />

1. Macfadyena unguis-cati (L.) A. H. Gentry,<br />

Brittonia 25: 236. 1973.<br />

Fig. 3. D; 45. A-C<br />

BASIONYM: Bignonia unguis-cati L.<br />

SYNONYMS: Batocydia unguis (L.) DC.<br />

Doxantha unguis-cati (L.) Miers<br />

Uña de gato, Liana uñada, Cat-claw<br />

Woody vine, which climbs by tendrils <strong>and</strong><br />

attains 10-15 m in length. Stems cylindrical,<br />

lenticellate, up to 6 cm in diameter; cross section<br />

<strong>of</strong> <strong>the</strong> mature stem with multilobed xylem, <strong>the</strong><br />

lobes alternating with radially arranged phloem<br />

tissue; nodes thickened; interpetiolar zone not<br />

gl<strong>and</strong>ular. Pseudostipules ovate, ca. 5 mm long.<br />

Leaves opposite, 2-foliolate, with a terminal<br />

tendril, trifid like a claw, generally <strong>of</strong> short<br />

duration; leaflets 6-16 × 1.2-7 cm, elliptical,<br />

oblong, or obovate, chartaceous or coriaceous,<br />

glabrous or with punctiform scales, <strong>the</strong> apex acute<br />

or acuminate, <strong>the</strong> base acute, rounded, or unequal,<br />

<strong>the</strong> margins undulate or rarely denticulate; upper<br />

surface dark, shiny, with sunken venation; lower<br />

surface light green, dull, with prominent venation;<br />

6. MANSOA<br />

Distribution: In forests <strong>and</strong> coastal thickets.<br />

Also on Cayo Luis Peña, Vieques, St. Croix, <strong>and</strong><br />

St. Thomas; Cuba <strong>and</strong> Hispaniola.<br />

Public forests: Cambalache, Ceiba, Guajataca,<br />

Guánica, Río Abajo, Susúa, <strong>and</strong> Vega.<br />

petioles 1-4.5 cm long, petiolules 0.5-2.5 cm long,<br />

both glabrous. Flowers solitary or in pairs,<br />

axillary; pedicel ca. 2 cm long. Calyx green,<br />

campanulate, 12-16 mm long, with 5 unequal<br />

lobes; corolla brilliant yellow, infundibuliform,<br />

4-8 cm long, <strong>the</strong> limb 3-6 cm in diameter, with 5<br />

unequal lobes, rounded; stamens 4, didynamous,<br />

inserted; ovary covered with punctiform scales.<br />

Capsule linear, somewhat woody, brown, 25-95<br />

cm long; seeds numerous, 1-3.5 cm long, with 2<br />

membranaceous wings.<br />

Phenology: Flowering <strong>and</strong> fruiting throughout<br />

<strong>the</strong> year.<br />

Status: Native, very common.<br />

Distribution: In moist forests at middle <strong>and</strong><br />

lower elevations, from <strong>the</strong> coast to <strong>the</strong> Cordillera<br />

Central. Also on Culebra, Desecheo, Vieques, St.<br />

Croix, St. John, St. Thomas, Tortola, <strong>and</strong> <strong>Virgin</strong><br />

Gorda; throughout <strong>the</strong> Antilles <strong>and</strong> tropical <strong>and</strong><br />

subtropical America.<br />

Public forests: Cambalache, Ceiba, Desecheo,<br />

El Yunque, Guánica, Maricao, Río Abajo, Susúa,<br />

Toro Negro, <strong>and</strong> Vega.<br />

Lianas with tendrils. Stems cylindrical or subquadrangular; cross section <strong>of</strong> <strong>the</strong> mature stem with a<br />

cross <strong>of</strong> 4 or 8 dark arms formed by <strong>the</strong> phloem tissue; interpetiolar zone gl<strong>and</strong>ular or not; pseudostipules<br />

short, conical-obtuse. Leaves opposite, 2- or 3-foliolate, with a terminal trifid or simple tendril; petioles<br />

sometimes with an apical gl<strong>and</strong>ular zone. Flowers in terminal panicles or racemes or in axillary corymbs.<br />

Calyx crateriform, tubular, or campanulate, simple, truncate or 5-dentate; corolla white, lilac, purple,<br />

or reddish, zygomorphic, tubular-campanulate or infundibuliform; stamens 4, didynamous; ovary

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