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Vines and Climbing Plants of Puerto Rico and the Virgin Islands

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<strong>Vines</strong> <strong>and</strong> <strong>Climbing</strong> <strong>Plants</strong> <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s<br />

rounded, mucronate, <strong>the</strong> base obtuse or rounded,<br />

frequently unequal on <strong>the</strong> lateral leaflets, <strong>the</strong><br />

margins entire; upper surface strigulose; lower<br />

surface sericeous-pubescent, with prominent<br />

venation, ciliate; petiolules tomentose, 2-4 mm<br />

long; rachis ferruginous-tomentose, sulcate, 0.2-<br />

2 cm long; petioles ferruginous-tomentose,<br />

sulcate, 3-11.5 cm long; stipules lanceolate,<br />

pubescent, 3-4 mm long; stipels linear, 1-2 mm<br />

long. Inflorescences <strong>of</strong> axillary pseudoracemes,<br />

10-15 cm long, with 2 or 3 flowers grouped at <strong>the</strong><br />

nodes; peduncle tomentose or sericeous. Calyx<br />

campanulate, ca. 6 mm long, tomentose, <strong>the</strong> lobes<br />

linear; corolla pink or lavender, <strong>the</strong> st<strong>and</strong>ard ca.<br />

4 mm long, oblong or rounded, <strong>the</strong> wings longer<br />

21. VIGNA<br />

than <strong>the</strong> keel, <strong>the</strong> keel whitish. Legume flattened,<br />

linear, 4-7 × 0.3-0.4 cm, ferruginous-tomentose,<br />

with a curved beak at <strong>the</strong> apex, dehiscent by valves<br />

that twist back in a spiral. Seeds oblong, 4-5 mm<br />

long, reddish brown.<br />

Phenology: Collected in flower from October<br />

to January <strong>and</strong> in fruit from October to March.<br />

Status: Native, relatively common.<br />

Distribution: In disturbed areas such as<br />

roadsides, pastures, secondary forests, <strong>and</strong><br />

thickets, at middle <strong>and</strong> lower elevations. Also in<br />

<strong>the</strong> Greater Antilles, tropical continental America<br />

<strong>and</strong> introduced in tropical Africa.<br />

Public Forests: Carite, Guajataca, Maricao,<br />

Río Abajo, <strong>and</strong> Toro Negro.<br />

Herbaceous vines, twining, climbing, or creeping. Leaves alternate, trifoliolate; stipules <strong>of</strong> various<br />

forms, persistent; stipels subulate. Inflorescences <strong>of</strong> axillary or terminal pseudoracemes; bracts small.<br />

Calyx campanulate, bilabiate, with 5 lobes, two <strong>of</strong> which are almost completely united; corolla yellow,<br />

pink, or pale blue, <strong>the</strong> st<strong>and</strong>ard rounded, <strong>the</strong> wings unguiculate, with a narrow projection or spur at <strong>the</strong><br />

base, <strong>the</strong> keel recurved, as long as <strong>the</strong> wings; stamens 10, diadelphous, <strong>the</strong> an<strong>the</strong>rs <strong>of</strong> <strong>the</strong> same size;<br />

ovary sessile, with numerous ovules, <strong>the</strong> style curved, pubescent on <strong>the</strong> distal portion, <strong>the</strong> stigma<br />

lateral. Fruit an oblong or linear legume, flattened to cylindrical, dehiscent by valves that twist on<br />

opening; seeds flattened, quadrangular or almost reniform. A genus <strong>of</strong> about 100 species, <strong>the</strong> majority<br />

<strong>of</strong> tropical Africa <strong>and</strong> Asia.<br />

Key to <strong>the</strong> species <strong>of</strong> Vigna<br />

1a. Corolla uniformly brilliant yellow………………………………………………………..2<br />

1b. Corolla pink or violet-pink, changing to white or pale yellow with purplish lines .……..5<br />

2a. St<strong>and</strong>ard ca. 1.5 cm long; legumes 4-7 cm long…………………………………………..3<br />

2b. St<strong>and</strong>ard 6-10 mm long; legumes 1-4 cm long……………………………………………4<br />

3a. Leaflets chartaceous, ovate, acute at <strong>the</strong> apex; stipules sagittate at <strong>the</strong> base; legumes 4-7 cm long<br />

..............………………….....…………………………………………..………5. V. luteola<br />

3b. Leaflets fleshy, obovate, obtuse, or rounded at <strong>the</strong> apex; stipules truncate at <strong>the</strong> base; legumes 5-6<br />

cm long .........................……………………………………………………….....6. V. marina<br />

4a. St<strong>and</strong>ard ca. 6 mm long; legume 8-15 mm long, with 1-3 seeds …………………. V. hosei<br />

4b. St<strong>and</strong>ard ca. 1 cm long; legume 3-4 cm long, with 5-7 seeds……………....4. V. longifolia<br />

5a. St<strong>and</strong>ard < 1 cm long; legumes ascendant …………..…………………..7. V. peduncularis<br />

5b. St<strong>and</strong>ard 1.5-3 cm long; legumes sparse or pendulous…………………………………….6<br />

6a. Style spirally twisted……………………………………………………………………... 7<br />

6b. Style erect or sigmoid…………………………………………………………………….. 8

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