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Vines and Climbing Plants of Puerto Rico and the Virgin Islands

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230<br />

<strong>Vines</strong> <strong>and</strong> <strong>Climbing</strong> <strong>Plants</strong> <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s<br />

14 × 2-8 cm, in general convex, <strong>the</strong> venation<br />

pinnate, usually from near <strong>the</strong> base, <strong>the</strong> apex<br />

acuminate, <strong>the</strong> base rounded or obtuse, <strong>the</strong><br />

margins entire or undulate, revolute; upper surface<br />

dark green, shiny, with <strong>the</strong> venation generally<br />

sunken, translucent <strong>and</strong> yellowish; lower surface<br />

yellowish green, with <strong>the</strong> palmate venation<br />

prominent; petioles 5-7 mm long, thick, rugose.<br />

Inflorescences <strong>of</strong> terminal corymbiform racemes;<br />

pedicels thick, 10-12 mm long. Calyx<br />

campanulate, ca. 5 mm long, green, smooth, with<br />

five minute lobes; corolla campanulate, early<br />

deciduous, yellowish green, ca. 8 mm long, <strong>the</strong><br />

lobes ovate, ca. 5 mm long, reflexed; stamens 10,<br />

<strong>the</strong> filaments white, flattened, adnate to <strong>the</strong> base<br />

<strong>of</strong> <strong>the</strong> corolla, <strong>the</strong> an<strong>the</strong>rs golden yellow, <strong>the</strong> distal<br />

tube as long as <strong>the</strong> <strong>the</strong>ca; ovary inferior, <strong>the</strong> style<br />

simple, <strong>the</strong> stigma subcapitate. Fruit fleshy,<br />

globose, 5-8 mm in diameter, cardinal red, with<br />

<strong>the</strong> calyx persistent on <strong>the</strong> apical portion. Seeds<br />

numerous, ca. 1 mm long, foveate, obtusetrigonal.<br />

Phenology: Flowering <strong>and</strong> fruiting from June<br />

to March.<br />

Status: Native, ra<strong>the</strong>r common.<br />

Distribution: From high <strong>and</strong> moist regions<br />

along <strong>the</strong> Cordillera Central <strong>and</strong> <strong>the</strong> Sierra de<br />

Luquillo. Also throughout <strong>the</strong> Antilles.<br />

Public Forests: Carite, El Yunque, Guilarte,<br />

<strong>and</strong> Toro Negro.<br />

References: Luteyn, J.L. 1976. Notes on neotropical Vaccinieae (Ericaceae). 1. Gonocalyx-A genus<br />

new to Central America. Brittonia 28: 37-41. Luteyn, J.L. <strong>and</strong> R.L. Wilbur. 1977. New genera <strong>and</strong><br />

species <strong>of</strong> Ericaceae (Vaccinieae) from Costa Rica <strong>and</strong> Panama. Brittonia 29: 255-276. Luteyn, J.L.<br />

1996. Ericaceae. In: Harling, G. <strong>and</strong> L. Andersson (eds.) Flora <strong>of</strong> Ecuador. Vol. 54.<br />

23. Family EUPHORBIACEAE<br />

Key to <strong>the</strong> genera<br />

1a. Inflorescences <strong>of</strong> cymes with two trilobate foliaceous bracts at <strong>the</strong> base, forming a pseudanthium;<br />

staminate flowers with numerous stamens; leaves deeply trilobate (in our species)<br />

…………….………..………………………….………………..................…1. Dalechampia<br />

1b. Inflorescences <strong>of</strong> racemes, lacking foliaceous bracts; staminate flowers with 2-3 stamens; leaves<br />

simple (in our species) ..………………………..……..………….....................………. 2. Tragia<br />

1. DALECHAMPIA<br />

Twining vines or less frequently shrubs, monoecious, usually covered with stinging hairs, producing<br />

scarce watery latex when wounded. Leaves simple, alternate, 3-5-lobate, petiolate, with a pair <strong>of</strong> stipels<br />

at <strong>the</strong> base <strong>of</strong> <strong>the</strong> blade; stipules present. Flowers unisexual, actinomorphic, apetalous, clustered in<br />

bisexual cymes, with long peduncles <strong>and</strong> two foliaceous bracts, forming a pseudanthium. Staminate<br />

flowers in distal cymules; bracteoles with numerous resinous gl<strong>and</strong>s; calyx 4-6-valvate; stamens<br />

numerous, grouped on a short stipe to form a head, <strong>the</strong> an<strong>the</strong>rs short, opening along longitudinal<br />

sutures; pistillode absent. Pistillate flowers in basal cymules; calyx with 8-12 lobes with gl<strong>and</strong>ular<br />

margins; ovary superior, trilobate, tricarpellate, each carpel with a solitary ovule, <strong>the</strong> style simple, with<br />

a capitate or peltate stigma. Fruit a trilobate capsule, with one seed per locule. A genus <strong>of</strong> about 100<br />

species, mostly <strong>of</strong> <strong>the</strong> Neotropics.

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