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Vines and Climbing Plants of Puerto Rico and the Virgin Islands

Vines and Climbing Plants of Puerto Rico and the Virgin Islands

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<strong>Vines</strong> <strong>and</strong> <strong>Climbing</strong> <strong>Plants</strong> <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s 341<br />

33. Family PHYTOLACCACEAE<br />

Key to <strong>the</strong> genera<br />

1a. Fruits woody or subwoody…………………………………………………………….......…… 2<br />

1b. Fruits fleshy ……………………………………………………….……...............… 3. Trichostigma<br />

2a. Fruits indehiscent, with <strong>the</strong> sepals much longer than <strong>the</strong> fruit …………….…........ 1. Agdestis<br />

2b. Fruits capsular, with <strong>the</strong> sepals shorter than <strong>the</strong> fruit ……………………............ 2. Stegnosperma<br />

1. AGDESTIS<br />

A monotypic genus, characterized by <strong>the</strong> following species.<br />

1. Agdestis clematidea Moçiño & Sessé ex DC.,<br />

Syst. Nat. 1: 543. 1818.<br />

Fig. 138. A-C<br />

Twining vine, herbaceous, attainig 15 m in<br />

length. Stems angular, reddish, striate, puberulent.<br />

Leaves with a strong, disagreeable odor, alternate,<br />

chartaceous, ovate or broadly ovate, 3-9 × 3-7.5<br />

cm, <strong>the</strong> apex obtuse or short-acuminate,<br />

mucronate, <strong>the</strong> base deeply cordiform or hastate,<br />

<strong>the</strong> margins crenulate; upper surface dark green,<br />

dull, with minute dots; lower surface light green,<br />

dull, puberulent, with prominent venation;<br />

petioles 1.5-9 cm long, with a reddish tinge,<br />

sulcate; stipules absent. Flowers bisexual, white,<br />

in axillary panicles, 6-17 cm long; peduncles<br />

2. STEGNOSPERMA<br />

glabrous or puberulent; pedicels 1-2 mm long,<br />

with a minute green bracteole, lanceolate, at <strong>the</strong><br />

base. Sepals 4, white, 3-5 mm long, oblong or<br />

oblanceolate, with <strong>the</strong> parallel venation<br />

conspicuous; petals absent; stamens 13-20; ovary<br />

partially inferior, <strong>the</strong> style conical, with 4 recurved<br />

stigmatic branches; ovule solitary. Fruits<br />

coriaceous, indehiscent, turbinate, ca. 3 mm long,<br />

with <strong>the</strong> sepals persistent at <strong>the</strong> base. Seeds<br />

elliptical.<br />

Phenology: Flowering from May to December.<br />

Status: Exotic, cultivated <strong>and</strong> naturalized<br />

(according to Liogier, 1985), uncommon.<br />

Distribution: Occasional in our gardens.<br />

Native to Mexico, cultivated sporadically in <strong>the</strong><br />

Antilles <strong>and</strong> South America.<br />

Erect or sc<strong>and</strong>ent shrubs; glabrous. Leaves alternate, simple; stipules minute, deciduous. Flowers<br />

bisexual, in terminal racemes; bracts <strong>and</strong> bracteoles minute, persistent. Calyx <strong>of</strong> 5 sepals; corolla <strong>of</strong> 5<br />

deciduous petals; stamens 10; ovary superior, with 3-5 uniovulate carpels, <strong>the</strong> styles free, as numerous<br />

as <strong>the</strong> carpels. Fruit a globose capsule, dehiscent from <strong>the</strong> apex toward <strong>the</strong> base; seeds 1-5, globose. A<br />

genus <strong>of</strong> 3 species, <strong>of</strong> <strong>the</strong> Antilles <strong>and</strong> Central America.<br />

1. Stegnosperma cubense A. Rich. in Sagra, Hist.<br />

Nat. Cub. 10: 309. 1845.<br />

Fig. 138. D-H<br />

Erect, clambering, or creeping shrub, with<br />

numerous lateral branches, attainig 1.5-5 m in<br />

length. Stems cylindrical, glabrous, dark gray,<br />

with minute lenticels. Leaves alternate, coriaceous

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