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Vines and Climbing Plants of Puerto Rico and the Virgin Islands

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120<br />

<strong>Vines</strong> <strong>and</strong> <strong>Climbing</strong> <strong>Plants</strong> <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s<br />

section <strong>of</strong> <strong>the</strong> mature stem with a cross <strong>of</strong> 8 dark,<br />

marginal arms, formed by <strong>the</strong> phloem tissue;<br />

branches quadrangular, glabrous, pseudostipules<br />

absent. Leaves opposite, 2-foliolate, with a<br />

terminal tendril, simple, filiform, early deciduous;<br />

lateral leaflets 5.5-16 × 2.5-9 cm, ovate, oblong,<br />

or elliptical, chartaceous, <strong>the</strong> apex obtuse, acute,<br />

or acuminate, <strong>the</strong> base obtuse to subcordiform,<br />

<strong>the</strong> margins entire; upper surface glabrous; lower<br />

surface with punctiform scales, without domatia;<br />

petioles <strong>and</strong> petiolules quadrangular, pubescent<br />

or glabrous, <strong>the</strong> petiole 0.9-4.5 cm long, <strong>the</strong><br />

petiolules 0.8-4 cm long. Flowers fragrant,<br />

produced in axillary or terminal panicles. Calyx<br />

yellowish, crateriform, simple, 4-10 mm long,<br />

pubescent or with punctiform scales, <strong>the</strong> apex<br />

truncate or denticulate; corolla zygomorphic,<br />

1. Distictis lactiflora (Vahl) DC., Prodr. 9: 191.<br />

1845.<br />

Fig. 3. A; 44. E-H<br />

BASIONYM: Bignonia lactiflora Vahl<br />

SYNONYM: Macrodiscus lactiflorus (Vahl) Bureau<br />

Pega palo, Liana fragante, Viuda<br />

Woody vine that climbs by tendrils with<br />

adhesive discs <strong>and</strong> attains 5 m or more in length.<br />

Mature stems cylindrical with <strong>the</strong> bark light<br />

brown, shredded; cross section with peripheral<br />

phloem forming a cross <strong>of</strong> 4 arms. Branches<br />

quadrangular, striate, puberulous, with thickened<br />

nodes; pseudostipules foliaceous, 3-4 mm long,<br />

deciduous. Leaves opposite, 2-foliolate, with a<br />

terminal tendril with 3 branches, 2 <strong>of</strong> <strong>the</strong>m<br />

modified into an adhesive disc; leaflets 2.7-8 ×<br />

2-4.5 cm, oblong, elliptical to subrounded,<br />

coriaceous, with <strong>the</strong> reticulate venation<br />

conspicuous, <strong>the</strong> apex obtuse, acute, or acuminate,<br />

4. DISTICTIS<br />

tubular-campanulate, 2.5-7.5 cm long, <strong>the</strong> tube<br />

slightly compressed, narrow at <strong>the</strong> base, white,<br />

yellow inside, <strong>the</strong> lobes rounded, lilac, <strong>the</strong> throat<br />

yellow with violet lines; stamens inserted; ovary<br />

cylindrical, 2-3 mm long, without a disc at <strong>the</strong><br />

base. Capsules linear, 21-43 × 1.5-2.4 cm,<br />

prominent on <strong>the</strong> margins; seeds numerous,<br />

semicircular, membranaceous, 1.5-3.8 cm long,<br />

with two membranaceous wings.<br />

Phenology: Flowering throughout <strong>the</strong> year.<br />

Status: Native, common.<br />

Distribution: In river-bank forests, on <strong>the</strong><br />

edges <strong>of</strong> mangrove swamps, <strong>and</strong> in coastal forests.<br />

Also on St. Croix, St. John, <strong>and</strong> St. Thomas; Cuba,<br />

Hispaniola, <strong>the</strong> Lesser Antilles, <strong>and</strong> continental<br />

tropical America.<br />

Public forest: Guánica.<br />

Lianas with tendrils. Stems hexagonal or quadrangular, cross section <strong>of</strong> <strong>the</strong> mature stem with<br />

peripheral phloem forming a cross <strong>of</strong> 4 arms, or cylindrical <strong>and</strong> not forming a cross; interpetiolar zone<br />

not gl<strong>and</strong>ular; pseudostipules present. Leaves opposite, 2-foliolate, frequently with a terminal trifid<br />

tendril. Flowers in terminal racemes or panicles. Calyx crateriform, simple, more or less truncate,<br />

usually gl<strong>and</strong>ular near <strong>the</strong> apex; corolla white, cream-colored, or red-violet, tubular-campanulate or<br />

broadly tubular, pubescent outside; stamens 4, didynamous; ovary superior, oblong, with 2 locules <strong>and</strong><br />

an annular-patelliform disc at <strong>the</strong> base. Fruit a convex or biconvex capsule, elliptic-oblong, with <strong>the</strong><br />

valves parallel to <strong>the</strong> septum; seeds compressed, with elongate irregular protuberances, in 2 rows. A<br />

genus <strong>of</strong> 6 species distributed from Mexico to Brazil, including <strong>the</strong> Antilles.<br />

<strong>the</strong> base rounded or acute, unequal; upper surface<br />

shiny, glabrous, puberulous, or minutely lepidote;<br />

lower surface pale green, dull, covered with<br />

minute transparent scales, venation prominent;<br />

petioles 1-2 cm long, <strong>the</strong> petiolules 5-10 mm long,<br />

both with 2 rows <strong>of</strong> crimped hairs. Flowers<br />

fragrant, in pendulous, terminal racemes or<br />

panicles, 10-30 cm long; peduncles <strong>and</strong> pedicels<br />

pubescent. Calyx simple, green, crateriform, 5-7<br />

mm long, truncate at <strong>the</strong> apex or minutely<br />

denticulate; corolla white, infundibuliform, 2.5-<br />

5.5 cm long, <strong>the</strong> tube yellow inside, <strong>the</strong> lobes 5,<br />

rounded, ca. 1.5 cm long. Capsule woody,<br />

elliptical, 6.5-13 × 1.7-3.5 cm, somewhat<br />

compressed or subcylindrical, with <strong>the</strong> apex <strong>and</strong><br />

base acute; seeds numerous, compressed, ca. 2.5<br />

cm long, <strong>the</strong> central circular body ca. 7 mm in<br />

diameter, with a surrounding marginal hyaline<br />

wing.

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