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Vines and Climbing Plants of Puerto Rico and the Virgin Islands

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<strong>Vines</strong> <strong>and</strong> <strong>Climbing</strong> <strong>Plants</strong> <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s 419<br />

25-60 × 0.4-1.0 cm, pleated or almost flat, with<br />

three main parallel veins, subrigid, retrorsely<br />

scabrous on <strong>the</strong> veins <strong>and</strong> <strong>the</strong> margins, <strong>the</strong> apex<br />

acuminate or obtuse; leaf sheath as long as <strong>the</strong><br />

internode, scabrous on <strong>the</strong> angles. Panicles<br />

solitary, distal or axillary, pyramidal, <strong>the</strong> distal<br />

ones 6-14 cm long, with a short leafy bract at <strong>the</strong><br />

base; axes minutely pubescent or glabrous;<br />

bracteoles filiform. Staminate <strong>and</strong> pistillate<br />

spikelets 4-6 mm long; sterile scales 4-5, ovatecircular,<br />

brown or reddish brown, finely ciliate<br />

on <strong>the</strong> margins, <strong>the</strong> apex apiculate. Achenes<br />

ovoid-globose or ovoid, 2.6-3.3 mm long, white,<br />

shiny or with a purple tinge; hypogynium<br />

trilobate, 1-1.8 mm long, whitish, <strong>the</strong> lobes<br />

semicircular, with <strong>the</strong> margins reflexed <strong>and</strong><br />

appressed to <strong>the</strong> achene.<br />

Phenology: Flowering <strong>and</strong> fruiting throughout<br />

<strong>the</strong> year.<br />

Status: Apparently endemic to <strong>Puerto</strong> <strong>Rico</strong>.<br />

Locally very common.<br />

Distribution: In moist or rain forests <strong>of</strong> <strong>the</strong><br />

cordilleras. Reported for Cuba, but no specimens<br />

seen or found.<br />

Public Forests: Carite, El Yunque, Guilarte,<br />

Maricao, <strong>and</strong> Toro Negro.<br />

2. Scleria scindens Nees ex Kunth, Enum. Pl. 2:<br />

343. 1837.<br />

Fig. 173. C-E<br />

Razor grass<br />

Perennial herb, forming large colonies, erect<br />

or climbing, attainig 3.5 m in length. Culms<br />

acutely triquetrous, 3-8 mm wide retrorsely <strong>and</strong><br />

antrorsely scabrid on <strong>the</strong> corners, strigulose on<br />

<strong>the</strong> distal portion. Leaves alternate, linear, 30-90<br />

× 0.5-1.5 cm, almost flat or pleated, with five<br />

parallel main veins, subrigid, retrorsely scabrid<br />

on <strong>the</strong> veins <strong>and</strong> <strong>the</strong> margins; leaf sheath scabrid<br />

on <strong>the</strong> angles. Panicles solitary, terminal,<br />

pyramidal, dense, 3-8 cm wide, sometimes with<br />

one or two small panicles at <strong>the</strong> base; bracts<br />

usually 3 on <strong>the</strong> terminal panicles. Staminate <strong>and</strong><br />

pistillate spikelets intermingled, <strong>of</strong> <strong>the</strong> same size;<br />

sterile scales 2-3, 3-4 mm long, light brown, ciliate<br />

on <strong>the</strong> margins, apiculate at <strong>the</strong> apex. Achenes<br />

globose, 2.3-2.6 mm long, apiculate, white, shiny,<br />

sometimes with a purple tinge; hypogynium<br />

trilobate, 1-1.7 mm long, <strong>the</strong> lobes semicircular,<br />

slightly revolute, divergent (not appressed to <strong>the</strong><br />

achene).<br />

Phenology: Collected in flower <strong>and</strong> fruit from<br />

February to August.<br />

Status: Native, locally common.<br />

Distribution: In moist or semi-moist disturbed<br />

areas in <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s (St. John, Tortola, <strong>and</strong><br />

<strong>Virgin</strong> Gorda). Also in Cuba, Hispaniola, <strong>and</strong> <strong>the</strong><br />

Lesser Antilles.<br />

3. Scleria secans (L.) Urb., Symb. Antill. 2(2):<br />

169. 1900.<br />

Fig. 173. F-G<br />

BASIONYM: Schoenus secans L.<br />

Perennial herb, rhizomatous, climbing,<br />

attainig 5-10 m in length. Culms branched in <strong>the</strong><br />

distal portion, acutely triquetrous, retrorsely<br />

spinulose on <strong>the</strong> corners, distally hirsute. Leaves<br />

alternate, linear, 10-40 × 0.2-0.7 cm, nearly flat<br />

or pleated, attenuate at <strong>the</strong> apex, <strong>the</strong> upper surface<br />

hirsute at <strong>the</strong> base, <strong>the</strong> lower surface sparsely<br />

hirsute to glabrous, retrorsely scabrous along <strong>the</strong><br />

midvein <strong>and</strong> <strong>the</strong> margins; leaf sheath scaberulous<br />

on <strong>the</strong> angles. Inflorescences <strong>of</strong> few spikelets,<br />

composed <strong>of</strong> a terminal panicle <strong>and</strong> 1-3 axillary<br />

panicles that are borne from <strong>the</strong> ultimate leaf;<br />

terminal panicle 2.5-6 cm long; <strong>the</strong> lateral ones<br />

shorter. Staminate spikelets shorter than <strong>the</strong><br />

pistillate ones; sterile scales 4, ovate or ovatelanceolate,<br />

2.5-6 mm long, light or dark brown,<br />

reddish or blackish on <strong>the</strong> margins. Achenes ovoid<br />

or oblong-ovoid, rounded at <strong>the</strong> apex, appressedpubescent,<br />

2.8-4 mm long, white, sometimes with<br />

greenish spots; hypogynium discoid, obscurely<br />

trilobate, 0.3-1 mm long, <strong>the</strong> lobes entire, reflexed.<br />

Phenology: Collected in flower <strong>and</strong> fruit from<br />

October to May.<br />

Status: Native, uncommon.<br />

Distribution: In thickets, along trails, <strong>and</strong> at<br />

<strong>the</strong> margin <strong>of</strong> moist forests along <strong>the</strong> Cordillera<br />

Central <strong>and</strong> <strong>the</strong> Sierra de Luquillo. Also in <strong>the</strong><br />

Antilles, Mexico, Central America, <strong>and</strong> South<br />

America.<br />

Public Forests: Carite <strong>and</strong> El Yunque.<br />

Reference: Strong, M. T. <strong>and</strong> P. Acevedo-Rodríguez. (in prep.) Cyperaceae. In: Acevedo-Rodríguez,<br />

P. & M.T. Strong (eds.) Monocots <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s. Contrib. U.S. National<br />

Herbarium.

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