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Vines and Climbing Plants of Puerto Rico and the Virgin Islands

Vines and Climbing Plants of Puerto Rico and the Virgin Islands

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<strong>Vines</strong> <strong>and</strong> <strong>Climbing</strong> <strong>Plants</strong> <strong>of</strong> <strong>Puerto</strong> <strong>Rico</strong> <strong>and</strong> <strong>the</strong> <strong>Virgin</strong> Isl<strong>and</strong>s 181<br />

or lavender, usually reddish at <strong>the</strong> base in <strong>the</strong><br />

center, infundibuliform or campanulate, 1.4-1.6<br />

cm long, <strong>the</strong> limb ca. 1 cm in diameter; stamens<br />

sagittate, white, two <strong>of</strong> <strong>the</strong>m shorter, not exserted;<br />

stigmas globose, white, not exserted. Capsule<br />

depressed-globose, brown, ca. 8 mm in diameter,<br />

pilose, with <strong>the</strong> style persistent; seeds 4 per fruit,<br />

3-5 mm long, glabrous, dark brown to black.<br />

Phenology: Flowering <strong>and</strong> fruiting throughout <strong>the</strong><br />

year.<br />

Status: Native, very common.<br />

Distribution: In grassl<strong>and</strong>s <strong>and</strong> pastures <strong>and</strong> on<br />

roadsides. Also on Icacos, Caja de Muerto,<br />

Desecheo, Mona, Vieques, St. Croix, St. John,<br />

St. Thomas, Tortola, <strong>and</strong> <strong>Virgin</strong> Gorda.<br />

Naturalized in tropical regions <strong>of</strong> <strong>the</strong> Old<br />

World.<br />

Public forests: Boquerón, Guánica, Mona,<br />

Piñones, <strong>and</strong> Río Abajo.<br />

Commentary: In <strong>the</strong> absence <strong>of</strong> <strong>the</strong> corolla, this<br />

species can be confused with I. tiliacea in<br />

individuals with relatively large <strong>and</strong> unlobed<br />

leaves. Never<strong>the</strong>less, I. triloba is distinguished<br />

by having pilose sepals, while those <strong>of</strong> I.<br />

tiliacea are glabrous.<br />

23. Ipomoea violacea L., Sp. Pl. 161. 1753.<br />

Fig. 1. B; 72. I-K<br />

SYNONYMS: Convolvulus tuba Schltdl.<br />

Calonyction tuba (Schltdl.) Colla<br />

Ipomoea tuba (Schltdl.) G. Don<br />

Ipomoea macrantha Roem. & Schult.<br />

Convolvulus gr<strong>and</strong>iflorus L.f.<br />

Operculina gr<strong>and</strong>iflora (L.f.) House, pro parte<br />

Bejuco de vaca, Flor de luna, Coast moon vine,<br />

Beach morning glory<br />

Slightly woody vine, twining, up to 15 m in<br />

length, with abundant milky latex. Stems<br />

cylindrical or subtriangular, grayish, with<br />

numerous lenticels usually arranged in lines; adult<br />

stems up to 3 cm in diameter, cross section with<br />

numerous alternating concentric rings <strong>of</strong> xylem<br />

<strong>and</strong> phloem. Leaves alternate; blades 9-15 × 7-11<br />

cm, glabrous, broadly ovate, sometimes trilobed,<br />

fleshy-coriaceous, usually involute, <strong>the</strong> apex acute<br />

or acuminate, mucronate, <strong>the</strong> base cordiform or<br />

deeply cordiform, <strong>the</strong> margins undulate; upper<br />

surface light green, with <strong>the</strong> venation yellowish;<br />

lower surface pale green, shiny, with prominent<br />

venation; petioles cylindrical, glabrous, yellowish<br />

green, longer than <strong>the</strong> blade, with a pair <strong>of</strong><br />

triangular gl<strong>and</strong>s at <strong>the</strong> base <strong>of</strong> <strong>the</strong> blade, swollen<br />

at <strong>the</strong> base. Flowers nocturnal, solitary or in<br />

simple dichasial cymes, axillary; peduncles<br />

cylindrical, up to 7 cm long, with two bracteoles<br />

in <strong>the</strong> upper middle portion. Calyx yellowish<br />

green, accrescent, <strong>the</strong> sepals unequal, 2-2.5 cm<br />

long, fleshy, concave, glabrous, <strong>the</strong> apex obtuse<br />

to rounded; corolla white, hypocrateriform,<br />

coriaceous, 5-7 cm long, <strong>the</strong> tube externally<br />

yellowish, internally with numerous transparent<br />

hairs <strong>and</strong> cardinal red fringes at <strong>the</strong> base, <strong>the</strong> limb<br />

6-7 cm in diameter, with 5 obtuse lobes; stamens<br />

<strong>and</strong> stigmas not exserted. Capsules globose or<br />

depressed-globose, yellowish brown, 2.5-3 cm in<br />

diameter, with accrescent sepals, persistent at <strong>the</strong><br />

base; seeds 4, obtusely angular, 1-1.2 cm long,<br />

brown, velvety, with a row <strong>of</strong> hairs along two<br />

angles.<br />

Phenology: Flowering throughout <strong>the</strong> year.<br />

Status: Native, common.<br />

Distribution: On <strong>the</strong> s<strong>and</strong>y coasts <strong>of</strong> <strong>the</strong> littoral<br />

zone. Also on Cayo Ratones, Culebra, Desecheo,<br />

Mona, Vieques, St. Croix, St. John, St. Thomas,<br />

<strong>and</strong> Tortola; <strong>the</strong> Antilles, <strong>the</strong> Bahamas, <strong>the</strong><br />

Cayman Isl<strong>and</strong>s, <strong>the</strong> United States (Florida), <strong>and</strong><br />

from Mexico to <strong>the</strong> Guianas. Introduced in <strong>the</strong><br />

tropics <strong>of</strong> <strong>the</strong> Old World.<br />

Public forests: Guánica, Mona, <strong>and</strong> Piñones.<br />

24. Ipomoea wrightii A. Gray, Syn. Fl. N. Amer.<br />

2(1): 213. 1878.<br />

Fig. 72. D-H<br />

SYNONYM: Ipomoea pulchella sensu Griseb.<br />

Herbaceous vine, annual, twining, climbing,<br />

or creeping, 3-4 m in length. Stems cylindrical,<br />

slender, glabrous, sometimes with minute<br />

spiniform projections. Leaves alternate, palmately<br />

compound; leaflets 5, lanceolate or oblonglanceolate,<br />

1-4 cm long, glabrous, chartaceous,<br />

<strong>the</strong> apex acute, <strong>the</strong> base acuminate, <strong>the</strong> margins<br />

entire; upper surface sparsely punctate, with <strong>the</strong><br />

midvein slightly prominent; lower surface pale<br />

green, with prominent venation, punctate or<br />

foveolate; petioles 2-6 cm long, slender, glabrous,<br />

with minute spiniform projections. Flowers<br />

usually solitary, axillary; peduncles 2-4 cm long,<br />

filiform, sinuate or spirally twisted; pedicels thick,<br />

claviform, 0.6-1.5 cm long, usually verrucose on

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