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the Symposium on Wheats for More Tropical Environments - cimmyt

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103hence, select <strong>for</strong> drought tolerance. Theexperiment c<strong>on</strong>sisted of 45 treatmentcombinati<strong>on</strong>s, five varieties and nineirrigati<strong>on</strong> treatments (Table 2).Due to wide variati<strong>on</strong>s betweenreplicates of different treatments, <str<strong>on</strong>g>the</str<strong>on</strong>g>rewas no significant difference between<str<strong>on</strong>g>the</str<strong>on</strong>g> varieties. Interacti<strong>on</strong> betweenirrigati<strong>on</strong> treatments and varieties wasalso not significant. Of all <str<strong>on</strong>g>the</str<strong>on</strong>g> varieties.<strong>on</strong>ly HD832-55 x eNO-JAR wasseriously affected when irrigati<strong>on</strong> waswithheld twice c<strong>on</strong>secutively at <str<strong>on</strong>g>the</str<strong>on</strong>g>jointing stage (T7). Missing twoirrigati<strong>on</strong>s at <str<strong>on</strong>g>the</str<strong>on</strong>g> flowering stage(T8) affected all of <str<strong>on</strong>g>the</str<strong>on</strong>g> varietiesc<strong>on</strong>siderably. Varying yield reducti<strong>on</strong>swere observed in all o<str<strong>on</strong>g>the</str<strong>on</strong>g>r treatments,but it appeared that <str<strong>on</strong>g>the</str<strong>on</strong>g> two mostcritical stages <strong>for</strong> moisture stress are T7and T8, Jointing and an<str<strong>on</strong>g>the</str<strong>on</strong>g>sis.Characteristics AffectingSelecti<strong>on</strong> <strong>for</strong> Heat and DroughtToleranceEarlinessAn understanding of a stress-tolerancemechanism is necessary in order tocorrectly suggest useful characteristicsby which a plant can tolerate stress.Under Nigerian c<strong>on</strong>diti<strong>on</strong>s, it appearsthat <str<strong>on</strong>g>the</str<strong>on</strong>g> main mechanism leading togood per<strong>for</strong>mance in wheat is escape oravoidance. The variety should grow andmature within <str<strong>on</strong>g>the</str<strong>on</strong>g> cold period, Le., withmaturities of 70 to 120 days. Thus.<strong>on</strong>ly early or medium-early lines!varieties will per<strong>for</strong>m well. This is alsotrue <strong>for</strong> drought tolerance. sinceearliness enhances water ec<strong>on</strong>omy.YieldIn additi<strong>on</strong> to earliness. superiority inyield under optimal c<strong>on</strong>diti<strong>on</strong>s isnecessary <strong>for</strong> both heat and droughttolerance. According to Blum (4)varieties with superior yield at optimallevels will also yield relatively wellunder sub-optimal levels.AdaptabiUtyWide adaptability is a necessary factor,since it is believed that. when stresstolerance is present in a variety. it maybe expressed as an unidentifiedcomp<strong>on</strong>ent of stability in per<strong>for</strong>manceover various envir<strong>on</strong>ments (4.11.17).Developmental synchr<strong>on</strong>yPlants should tiller uni<strong>for</strong>mly, holdmost of <str<strong>on</strong>g>the</str<strong>on</strong>g>ir leaves intact and matureuni<strong>for</strong>mly.Test weightThe grains of plants should have goodtest weight, which results from goodand uni<strong>for</strong>m grain filling.Spike fertUltyIncreased fertility of <str<strong>on</strong>g>the</str<strong>on</strong>g> spikes assuresthat most florets will be fertile understress c<strong>on</strong>diti<strong>on</strong>s, as well as at optimallevels.Table 2. Effects of variety and moisture stress <strong>on</strong> grain yield of bread wheat, Bakura,Nigeria, 1982·83Grain yield (kg/ha) according to i'rrigati<strong>on</strong> treatment!./Variety T1 T2 T3 T4 T5 T6 T7 T8 T9HD832-55 x CIANO-JAR 2747 1907 1720 1813 2087 1687 833 1587 2220Super X 2627 1893 1960 1667 1887 1987 1433 1147 1820(TF x CIAN067/Bluebird-CIANO)ZA 1840 2033 1860 1887 2527 1960 1733 1187 1867NP876-164 x Roch/CIANO"S"-Bluebird 2380 1960 1960 2020 1860 1600 1793 1453 1720Pi62·Fr<strong>on</strong>d/Pi62-Mazoe x Mexipak 65 2293 2427 1867 1487 2427 1860 1667 1333 2120!,/ T1 = c<strong>on</strong>trol, 19 irrigati<strong>on</strong>s; T2 = <strong>on</strong>e irrigati<strong>on</strong> withheld at tillering; T3 = <strong>on</strong>e withheld at jointing; T4 = <strong>on</strong>e withheldat flowering; T5 = <strong>on</strong>e withheld at grain filling; T6 = two irrigati<strong>on</strong>s withheld at tillering; T7 = two withheld at jointing;T8 = two withheld at flowering; T9 = two withheld at grain fillingSource: Aremu and Orakwe (2)

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