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__ Maturation 500 o_ Maturation<br />

_ 600 E<br />

r-- I _ --- 300<br />

400 -'=<br />

447.1 507.5 504.7 Mean .=,,_200 339 7 425 6 405.7 Mean<br />

• •<br />

200 30 21 19 N _= 100 37 27 20 N<br />

0<br />

_t Pupation o_ Pupation<br />

600 500<br />

500 ]....._ I --- 400<br />

40o I Ij F-__] , 3oo I 1 w<br />

300 El, _ 200 '<br />

._ 200 342.6 396.6 406.3 Mean 265.3 339.1 329.6 I_ean<br />

100 29 21 19 N _= 100 32 27 20 II J<br />

0 , , , 0 ' '<br />

A B C A B C<br />

Locationof Q. punctatella Locationof Q. punctatella<br />

Figure lO.---Body size of three different regional Q. punctatella populations reared on beeches in the Tohoku <strong>Research</strong><br />

Center of the F<strong>or</strong>estry and F<strong>or</strong>est Products <strong>Research</strong> Institute in M<strong>or</strong>ioka. Weight of larvae at maturation and at<br />

pupation are indicated f<strong>or</strong> each sex. Bar means maximum and minimum value, and upper and lower sides of box are<br />

one standard err<strong>or</strong>.<br />

Tests of Two Hypotheses in Field Populations<br />

Larvae from all of the regions were reared on beech saplings in each of the respective regions. Plant quality influenced<br />

surviv<strong>or</strong>ship in each region. Although great differences in surviv<strong>or</strong>ship between regions were observed, little difference<br />

was found within a region (Table 1). Approximately half of the larvae reared on beech saplings in Sites B and C<br />

survived to maturity, whereas surviv<strong>or</strong>ship of all those reared on Site A beech was between 3.2% and 6.5%. The influence of<br />

Q, p,nctawlla population quality in each region on body size was clearly evident. Irrespective of the location of beech<br />

saplings on which the larvae were reared, the <strong>or</strong>der of average body size among the regional populations was the same: B >__<br />

C 2 A (Table 2). In particular, body size of A population larvae was significantly smaller than that of larvae from the other<br />

Table !.--Surviv<strong>or</strong>ship of Q. punctatella larvae from each region on beech in that region.<br />

78<br />

Q. punctatella<br />

A B C<br />

A 3.6% 3.2% 6.5%<br />

(6/165) (5/155) (10/154)<br />

E crenata B 48.9% 55.0% 53.9%<br />

(44/90) (33/60) (41/76)<br />

C 55.8% 52.9%<br />

(72/129) (36/68)<br />

(No. of matured larvae / No. of hatched larvae)

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