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marker-assisted selection in wheat - ictsd

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436Marker-<strong>assisted</strong> <strong>selection</strong> – Current status and future perspectives <strong>in</strong> crops, livestock, forestry and fishered to farmers. Although documentationwas limited, the current impact seemedsmall while the future impact was likely tobe far more substantial.Priyadarshan (11) <strong>in</strong>dicated that biotechnologyresearch had been supportedactively for over 17 years <strong>in</strong> India, butwas doubtful about the impact on varietiesreleased to farmers. He believed thatresearch on MAS and other biotechnologieshad rema<strong>in</strong>ed largely <strong>in</strong> journal articlesand had not significantly boosted conventionalplant breed<strong>in</strong>g efforts on the ground.Kirti (12) lamented that there was no comprehensivedocumentation regard<strong>in</strong>g thesuccessful use of MAS for breed<strong>in</strong>g newcrop varieties or develop<strong>in</strong>g breed<strong>in</strong>g material,as this <strong>in</strong>formation would be importantfor evaluat<strong>in</strong>g the technology. Collard (45),while not<strong>in</strong>g that MAS had been successful<strong>in</strong> cereal crops <strong>in</strong> his country, Australia,said he was not aware of many examples ofMAS-derived cultivars grown <strong>in</strong> Australiadespite the wealth of publications fromAustralian <strong>in</strong>stitutions on the technology.Sridhar (48) suggested that, <strong>in</strong> India, mostproducts of MAS are still <strong>in</strong> the hands ofresearch <strong>in</strong>stitutions undergo<strong>in</strong>g evaluation.He suggested that MAS products require a“fast track” evaluation system to expeditethe release of promis<strong>in</strong>g germplasm.Accord<strong>in</strong>g to Makkar (52), success <strong>in</strong>demonstrat<strong>in</strong>g genetic ga<strong>in</strong> <strong>in</strong> the laboratorydid not always equate with successunder field conditions. However, some realsuccesses were reported, <strong>in</strong>clud<strong>in</strong>g transferof important resistance genes <strong>in</strong>to adaptedrice germplasm for Indian farmers (Sridhar,35 and 54), <strong>in</strong>dicat<strong>in</strong>g that more successesmight be <strong>in</strong> the pipel<strong>in</strong>e. Williams (51) saidthat molecular <strong>marker</strong>s had been used for atleast five years <strong>in</strong> Australia <strong>in</strong> some <strong>wheat</strong>and barley improvement programmes andthat “it is likely that <strong>in</strong> Australia all breed<strong>in</strong>gprogrammes with <strong>in</strong>dustry fund<strong>in</strong>g andprobably also the private breed<strong>in</strong>g companiesare currently us<strong>in</strong>g MAS to someextent”. However, the potential of the newtechnology has to be weighed aga<strong>in</strong>st thesuccess achieved us<strong>in</strong>g traditional methods.Acikgoz (66) po<strong>in</strong>ted out that the Turkishrice cultivar Tokak was still be<strong>in</strong>g solddespite hav<strong>in</strong>g been released <strong>in</strong> 1937, andquestioned how much impact populationgenetics studies, popular 20–30 years ago,had on cultivar development, let alone theimpact of biotechnology applications.Buijs (58) mentioned tissue culture, onceregarded as a modern, relatively expensivetechnology, which is now relatively <strong>in</strong>expensiveand widely used <strong>in</strong> develop<strong>in</strong>gcountries. It will only be known retrospectivelywhether MAS evolves similarlyto become a standard tool of the plant andanimal breeder <strong>in</strong> develop<strong>in</strong>g countries.Intellectual property rights issuesSome participants felt that <strong>in</strong>tellectualproperty rights (IPRs) were an importantconstra<strong>in</strong>t to development and uptakeof MAS <strong>in</strong> the develop<strong>in</strong>g world. Corva(29) raised the issue of the use of licensedgenomic technology by public <strong>in</strong>stitutions<strong>in</strong> develop<strong>in</strong>g countries, mention<strong>in</strong>g thatmany useful cattle <strong>marker</strong>s were becom<strong>in</strong>gavailable, but which were patented, andthat there was therefore a demand for practical<strong>in</strong>formation about IPRs and violationof IPRs. Weller (30) po<strong>in</strong>ted out that patentsare only valid <strong>in</strong> the country wherethey are granted, that research tends to beexempted from patent restrictions and thatthere can be long delays between fil<strong>in</strong>g ofpatent claims and their eventual grant<strong>in</strong>g.Saravanan (31) argued strongly for thefreedom of researchers to use patented biotechnologytools. Storlie (32) argued thatfarmers <strong>in</strong> the develop<strong>in</strong>g world should

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