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Lost River - Karst Information Portal

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Figure 8. Skull of the dire wolf, Canis dirus,<br />

from the Microfauna Room, Megenity Peccary<br />

Cave, Crawford County, Indiana. This wolf<br />

suffered a dislocated rear leg, likely rendering<br />

him susceptible to entrapment. Fred Lewis photo.<br />

Scale in centimeters.<br />

shrew (Sorex arcticus), northern bog lemming<br />

(Mictomys borealis), and yellow-cheeked vole<br />

(Microtus xanthognathus), was dated at 14,125<br />

BP, long after peccary accumulation had ceased.<br />

Other extinct taxa are represented by several<br />

osteoderms of the beautiful armadillo (Dasypus<br />

bellus), a tooth portion of a tapir (Tapirus sp),<br />

five teeth of the long-nosed peccary (Mylohyus<br />

nasutus), and one disintegrated tooth of a<br />

horse. Additional extralimital taxa include<br />

heather vole (Phenacomys intermedius), boreal<br />

red-backed vole (Clethrionomys gapperi), and<br />

snowshoe hare (Lepus americanus). More<br />

recent accumulations are from the activities<br />

of woodrats, wandering carnivores (scat),<br />

and humans. After the grueling 24 years of<br />

excavation have been completed will come the<br />

great reward of cataloguing and analysis. The<br />

occurrence of any bone recovered in the cave<br />

can be plotted, revealing concentrations and<br />

suggesting patterns of selection, accumulation,<br />

and change through time.<br />

A chapter on the origins of and change in<br />

Indiana’s vertebrate fauna, with descriptions of<br />

cave fossil occurrences (Richards and Whitaker,<br />

1997), was included in The Natural Heritage of<br />

Indiana.<br />

The biggest surprise of Indiana’s karst<br />

faunas was the 1996 discovery of a bone-rich<br />

sinkhole deposit in Pipe Creek Junior Quarry,<br />

<strong>Karst</strong> Vertebrate Paleobiology in Indiana<br />

Grant County, containing the remains of<br />

extinct rhinoceros, camel, llama, giant land<br />

tortoise, and others among abundant bones<br />

of turtles, frogs, snakes, and plant macrofossils<br />

(Farlow, et al., 2001). Initially formed during<br />

early Pliocene and or late Miocene times as a<br />

collapsed cave roof, the passage became plugged<br />

with sediment, forming a sinkhole pond at the<br />

surface. An aquatic biota flourished, leaving<br />

their remains in the dark organic pond fill,<br />

supplemented by the bones and teeth of larger<br />

animals (Figure 9). The 5 million year old<br />

northern Indiana sinkhole remains were later<br />

overridden by the ice of numerous glaciations<br />

until discovery. The biota, under study by<br />

numerous specialists, is among the few faunas of<br />

its age in the interior of eastern North America<br />

(Farlow, et al., 2001).<br />

Figure 9. Artist’s rendering of the Pipe Creek<br />

Sinkhole Biota, Grant County, northern Indiana.<br />

Some 5 million years ago, this karst pond supported<br />

a rich biota, including rhinoceros (Teleoceras sp.),<br />

bone-crushing dog (Borophagus sp.), bear<br />

(Plionarctos edensis) and a large species of peccary.<br />

Artist Karen Carr, Courtesy Indiana State<br />

Museum and Historic Sites.<br />

205

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