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Vegetation Classification and Mapping Project Report - USGS

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<strong>USGS</strong>-NPS <strong>Vegetation</strong> <strong>Mapping</strong> Program<br />

Colonial National Historical Park<br />

companion species. Other woody species include Persea palustris (swamp bay), Magnolia<br />

virginiana (sweetbay), Salix caroliniana (coastal plain willow), Vaccinium formosum (southern<br />

blueberry), Smilax walteri (coral greenbrier), <strong>and</strong> saplings of Acer rubrum (red maple), Pinus<br />

taeda (loblolly pine), Taxodium distichum (bald cypress), <strong>and</strong> Liquidambar styraciflua<br />

(sweetgum). Herbaceous composition is diverse <strong>and</strong> varied, with many species 'recruited' from<br />

adjacent marshes <strong>and</strong> swamps. Thelypteris palustris var. pubescens (eastern marsh fern) is highly<br />

characteristic <strong>and</strong> appears to reach its nodal distribution in this l<strong>and</strong>scape in this community.<br />

Other important species can include Juncus roemerianus (needlegrass rush), Cladium mariscus<br />

ssp. jamaicense (Jamaica swamp sawgrass), <strong>and</strong> Decodon verticillatus (swamp loosestrife).<br />

Murdannia keisak (wartremoving herb) is an aggressive alien weed.<br />

In Maryl<strong>and</strong>, these tidal oligohaline shrubl<strong>and</strong>s are characterized by a low (0.5-6 m tall), open<br />

(40-60% cover) to moderately dense (60-80% cover) shrub canopy dominated by Morella<br />

cerifera (wax myrtle) with Rosa palustris (swamp rose) as a frequent associate. Morella cerifera<br />

(wax myrtle) is most constant <strong>and</strong> typically occupies 25 to 50% of the shrub stratum. In general,<br />

Rosa palustris (swamp rose) is slightly less abundant (10-25% mean cover) than Morella<br />

cerifera (wax myrtle); however, it can occasionally be dominant (>75% cover) in some st<strong>and</strong>s.<br />

Other less frequent taxa in the shrub stratum include Acer rubrum (red maple), Viburnum<br />

recognitum (southern arrowwood), Baccharis halimifolia (eastern baccharis), <strong>and</strong> dense vine<br />

growth of Toxicodendron radicans (eastern poison ivy) <strong>and</strong> Mikania sc<strong>and</strong>ens (climbing<br />

hempvine). St<strong>and</strong>s along ecotones or in less frequently flooded situations may contain emergent<br />

individuals of Pinus taeda (loblolly pine), Liquidambar styraciflua (sweetgum), Fraxinus<br />

pennsylvanica (green ash), Fraxinus profunda (pumpkin ash), Nyssa biflora (swamp tupelo), <strong>and</strong><br />

Juniperus virginiana var. virginiana (eastern redcedar) in a scattered <strong>and</strong> very sparse (0-5%<br />

cover) overstory. Species richness in the herbaceous layer is very high <strong>and</strong> indicative of slight<br />

oligohaline conditions, species recruitment from adjacent communities, <strong>and</strong> considerable<br />

microtopographic variation. Osmunda regalis var. spectabilis (royal fern) is consistent <strong>and</strong><br />

dominant in the herbaceous layer, with Thelypteris palustris var. pubescens (eastern marsh fern)<br />

in close association although not as abundant. Associated species include species characteristic<br />

of freshwater marshes, such as Leersia oryzoides (rice cutgrass), Polygonum punctatum (dotted<br />

smartweed), <strong>and</strong> Pelt<strong>and</strong>ra virginica (green arrow arum), <strong>and</strong> species more tolerant of<br />

mesohaline conditions, such as Hibiscus moscheutos ssp. moscheutos (crimsoneyed rosemallow)<br />

<strong>and</strong> Carex hormathodes (marsh straw sedge). Other characteristic herbs include Apios americana<br />

(groundnut), Cuscuta gronovii (scaldweed), Sium suave (hemlock waterparsnip), Polygonum<br />

arifolium (halberdleaf tearthumb), Ptilimnium capillaceum (herbwilliam), Cinna arundinacea<br />

(sweet woodreed), Thalictrum pubescens (king of the meadow), Typha latifolia (broadleaf<br />

cattail), Cicuta maculata (spotted water hemlock), <strong>and</strong> Boehmeria cylindrica (smallspike false<br />

nettle).<br />

Most Abundant Species:<br />

Stratum Lifeform Species<br />

Tall shrub/sapling Broad-leaved deciduous shrub Rosa palustris (swamp rose),<br />

Salix caroliniana (coastal plain willow)<br />

Tall shrub/sapling Broad-leaved evergreen shrub Morella cerifera (wax myrtle)<br />

Characteristic Species: Morella cerifera (wax myrtle), Rosa palustris (swamp rose), Salix<br />

caroliniana (coastal plain willow), Thelypteris palustris var. pubescens (eastern marsh fern),<br />

Toxicodendron radicans ssp. radicans (eastern poison ivy).<br />

277

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