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Vegetation Classification and Mapping Project Report - USGS

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<strong>USGS</strong>-NPS <strong>Vegetation</strong> <strong>Mapping</strong> Program<br />

Colonial National Historical Park<br />

Heritage exemplary natural community occurrence because of its large size <strong>and</strong> overall high<br />

quality. Areas representing the other 11 natural communities mapped in Colonial National<br />

Historical Park do not meet the criteria to be considered Natural Heritage exemplary natural<br />

community occurrences, but these native plant associations are an important part of the park’s<br />

natural resources <strong>and</strong> should be targets for conservation <strong>and</strong> management.<br />

Invasive nonnative plant species are one of the main threats to the native vegetation associations<br />

at Colonial National Historical Park. Thirty-eight nonnative plant species were noted in the<br />

quantitative plots <strong>and</strong> accuracy assessment observations. The fertile, eroding soils of Coastal<br />

Plain Dry Calcareous Forest supported the greatest diversity of nonnative plant species, where 16<br />

nonnative species were noted. Of the other map classes, wetl<strong>and</strong>s <strong>and</strong> transitional vegetation had<br />

the highest cover by invasive, nonnative species. The most common <strong>and</strong> problematic species<br />

include Japanese honeysuckle (Lonicera japonica), Nepalese browntop (Microstegium<br />

vimineum), <strong>and</strong> common reed (Phragmites australis). Japanese honeysuckle <strong>and</strong> Nepalese<br />

browntop are particularly troublesome because of their shade tolerance <strong>and</strong> aggressive growth<br />

habits. These species can be opportunistic invaders of the older, more intact forest communities,<br />

getting a foothold where roads, trails, tip-up mounds, downfalls, <strong>and</strong> other gap-disturbances have<br />

disturbed mineral soil. Once established, colonies are able to more easily exp<strong>and</strong> or spread into<br />

nearby microhabitats. Common reed is an aggressive, invasive perennial grass that invades tidal<br />

wetl<strong>and</strong>s.<br />

Japanese honeysuckle is especially destructive to native vegetation because of its rapid, twining<br />

growth <strong>and</strong> dense, semi-evergreen foliage that shades out competitors. The vines frequently<br />

strangle shrubs <strong>and</strong> tree saplings, <strong>and</strong> over-grow more delicate herbs in a variety of settings.<br />

Nepalese browntop is the fastest spreading, most problematic exotic everywhere in Virginia<br />

today. Within the past two decades, it has abundantly invaded moist, openly shaded habitats<br />

throughout the mid-Atlantic region, forming monospecific carpets of tangled culms that tend to<br />

crowd out competing herbaceous species (Tu 2000). A recent study strongly suggests that<br />

Nepalese browntop responds to forest canopy disturbances with a sudden increase in biomass<br />

that impedes woody regeneration <strong>and</strong> lowers overall species diversity <strong>and</strong> stem densities (Oswalt<br />

et al. 2007). Other studies have demonstrated that once established, Nepalese browntop overruns<br />

native herbaceous competitors <strong>and</strong> leads to dramatic declines of herb richness within a few<br />

years (Barden 1987, Hunt <strong>and</strong> Zaremba 1992). Common reed (Phragmites australis) is an<br />

aggressive wetl<strong>and</strong> invader that out-competes native plant species, eliminates diverse wetl<strong>and</strong><br />

plant communities, <strong>and</strong> provides little food or shelter for wildlife. It grows in dense singlespecies<br />

st<strong>and</strong>s <strong>and</strong> forms a thick mat of roots <strong>and</strong> rhizomes, spreading by seed <strong>and</strong> rhizomes.<br />

Common reed becomes established in areas where soil is exposed through natural (storms, wave<br />

action) or anthropogenic disturbances (construction, ditching).<br />

Other highly invasive, nonnative species noted in the park include tree of heaven (Ailanthus<br />

altissima), autumn olive (Elaeagnus umbellata var. parvifolia), <strong>and</strong> Chinese lespedeza<br />

(Lespedeza cuneata). Species noted that are considered moderately invasive by the Virginia<br />

Department of Conservation <strong>and</strong> Recreation (VADCR 2003) include English Ivy (Hedera helix),<br />

golden bamboo (Phyllostachys aurea), princesstree (Paulownia tomentosa)Canada bluegrass<br />

(Poa compressa), rough bluegrass (Poa trivialis), white poplar (Populus alba), curley dock<br />

(Rumex crispus), <strong>and</strong> Chinese wisteria (Wisteria sinensis). Species considered occasionally<br />

invasive include orchardgrass (Dactylis glomerata), <strong>and</strong> common periwinkle (Vinca minor).<br />

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