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Fisheries in the Southern Border Zone of Takamanda - Impact ...

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130 Sunderland-Groves et al.<br />

In more recent years, gorilla groups <strong>in</strong> Cameroon-<br />

Nigeria border region <strong>the</strong> study area have become more<br />

fragmented and isolated as <strong>the</strong>ir habitat has succumbed to<br />

agricultural and o<strong>the</strong>r land development activities.<br />

Construction <strong>of</strong> <strong>the</strong> new road between <strong>the</strong> towns <strong>of</strong><br />

Mamfe and Akwaya, when completed, will cut directly<br />

between <strong>Takamanda</strong> and <strong>the</strong> Mone and Mbulu forests,<br />

prevent<strong>in</strong>g gorilla movement between <strong>the</strong> areas.<br />

Hunt<strong>in</strong>g and agricultural activities will have a similar<br />

effect on <strong>the</strong> chimpanzee populations <strong>of</strong> <strong>the</strong> area that, like<br />

<strong>the</strong> gorillas, are fragmented and conf<strong>in</strong>ed to highland<br />

areas. In 1997, <strong>the</strong> chimpanzees <strong>of</strong> eastern Nigeria and<br />

<strong>Takamanda</strong> were described as a dist<strong>in</strong>ct sub-species Pan<br />

troglodytes vellerosus (Gonder et al. 1997), and although<br />

<strong>the</strong>ir numbers <strong>in</strong> <strong>the</strong> wild have not been thoroughly<br />

<strong>in</strong>vestigated, it is thought that <strong>the</strong> populations are<br />

decl<strong>in</strong><strong>in</strong>g.<br />

The Cross River Gorilla Research Project<br />

(Cameroon) has <strong>in</strong>itiated conservation efforts to<br />

elim<strong>in</strong>ate hunt<strong>in</strong>g <strong>of</strong> <strong>the</strong>se two species <strong>in</strong> <strong>the</strong> region. The<br />

focus is on work<strong>in</strong>g with local communities and<br />

dissem<strong>in</strong>ation <strong>of</strong> educational materials <strong>in</strong> collaboration<br />

with <strong>the</strong> jo<strong>in</strong>t M<strong>in</strong>istry <strong>of</strong> Environment and Forests<br />

(MINEF)/GTZ project—Protection <strong>of</strong> <strong>the</strong> Forests<br />

Around Akwaya (PROFA).<br />

3 Study area<br />

The 675-km 2 <strong>Takamanda</strong> Forest Reserve is mostly<br />

covered by Atlantic evergreen forest, rang<strong>in</strong>g from 100 m<br />

to 1500 m above sea level (Figure 2 <strong>in</strong> Chapter 1). Much<br />

<strong>of</strong> <strong>the</strong> lowland forest <strong>in</strong> <strong>the</strong> sou<strong>the</strong>rn and central parts <strong>of</strong><br />

<strong>the</strong> Reserve is between 100 m and –400 m <strong>in</strong> elevation.<br />

The terra<strong>in</strong> is roll<strong>in</strong>g <strong>in</strong> <strong>the</strong> lowlands, but rises sharply to<br />

1500 m <strong>in</strong> <strong>the</strong> nor<strong>the</strong>rn part <strong>of</strong> <strong>the</strong> Reserve where slopes<br />

are extremely steep and areas <strong>of</strong> semi-deciduous forest,<br />

woody savannah (<strong>of</strong>ten degraded), and montane woody<br />

savannah with grasslands prevail between 1200 m and<br />

2200 m (Letouzey 1985). Around villages, <strong>the</strong> vegetation<br />

has been modified and is degraded evergreen lowland<br />

forest and farmbush (a mosaic <strong>of</strong> cultivation and fallow).<br />

The lowland forest is particularly diverse; it is thought to<br />

be part <strong>of</strong> a Pleistocene refugium. The forest formation is<br />

<strong>Takamanda</strong>: <strong>the</strong> Biodiversity <strong>of</strong> an African Ra<strong>in</strong>forest<br />

dist<strong>in</strong>ct as <strong>the</strong>re is a relative paucity <strong>of</strong> <strong>the</strong><br />

Caesalp<strong>in</strong>ioides, which are normally common <strong>in</strong> <strong>the</strong><br />

Atlantic coastal forest (see Sunderland et al. this volume<br />

for a more detailed discussion <strong>of</strong> <strong>the</strong> vegetation <strong>in</strong> <strong>the</strong><br />

area). 0<br />

In 1985, <strong>the</strong> human population density <strong>in</strong> <strong>the</strong><br />

<strong>Takamanda</strong> area was estimated at between 6 and 12<br />

people/km 2 (Letouzey 1985). In 1999 <strong>the</strong> approximate<br />

total human population, based on a complete census <strong>of</strong><br />

n<strong>in</strong>e villages, was estimated at 2,490 (Groves and<br />

Maisels 1999). A more recent socio-economic survey<br />

conducted by PROFAthat covered 43 villages with<strong>in</strong> and<br />

surround<strong>in</strong>g <strong>the</strong> <strong>Takamanda</strong> Forest Reserve, <strong>in</strong>clud<strong>in</strong>g 12<br />

villages on <strong>the</strong> Nigerian side <strong>of</strong> <strong>the</strong> border, estimated total<br />

human population at 15,707 (Schmidt-Soltau et al.<br />

2001). Socio-economic activities <strong>in</strong> <strong>the</strong> region revolve<br />

strongly around <strong>the</strong> forest and its resources, especially for<br />

<strong>the</strong> villages that are far from markets (Ifeka 1999,<br />

Sunderland et al. this volume), and people who live <strong>in</strong> <strong>the</strong><br />

most remote villages depend almost entirely on<br />

bushmeat for <strong>the</strong>ir prote<strong>in</strong> requirements. They also rely<br />

on <strong>the</strong> harvest and sale <strong>of</strong> non-timber forest products as<br />

<strong>the</strong>ir ma<strong>in</strong> source <strong>of</strong> cash, particularly Irv<strong>in</strong>gia<br />

gabonensis, or “bush mango,” ra<strong>the</strong>r than on cultivated<br />

crops or livestock.<br />

The area has been historically partially protected by<br />

lack <strong>of</strong> access, but this is chang<strong>in</strong>g. Accessibility to <strong>the</strong><br />

<strong>Takamanda</strong> Reserve from <strong>the</strong> Nigerian side is by foot<br />

only, but a new road from Mamfe (Figure 2 <strong>in</strong> Chapter 1)<br />

now allows access to with<strong>in</strong> 5 km <strong>of</strong> its limits from <strong>the</strong><br />

Cameroonian side. The Mone Forest Reserve is<br />

separated from <strong>the</strong> <strong>Takamanda</strong> reserve by a corridor <strong>of</strong><br />

ungazetted forest, which, at its narrowest, is about 7 km<br />

wide.<br />

Dur<strong>in</strong>g <strong>the</strong> 1998-1999 surveys, two ma<strong>in</strong> vegetation<br />

types were sampled: lowland forest and sub-montane<br />

forest. The study area comprised two lowland sites along<br />

<strong>the</strong> Makone and Oyi/Magbe rivers and three hill, or<br />

highland, areas—Matene, Obonyi, and Basho Hills—<br />

with<strong>in</strong> <strong>Takamanda</strong> Forest Reserve (Figure 2 <strong>in</strong> Chapter<br />

1). The Makone flows through <strong>the</strong> middle <strong>of</strong> <strong>the</strong> Reserve,<br />

and <strong>the</strong> Oyi forms part <strong>of</strong> <strong>the</strong> Reserve’s southwestern

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