24 Sunderland et al. Table 2. Summary <strong>of</strong> floristic data for plots <strong>in</strong> <strong>Takamanda</strong> Forest Reserve, Cameroon Site Plot # # <strong>of</strong> trees with species <strong>in</strong>clud<strong>in</strong>g Berl<strong>in</strong>ia bracteosa, B. craibiana, Afzelia pachyloba, A. bip<strong>in</strong>densis, Microberl<strong>in</strong>ia bisulcata, Erythrophloem ivorense, Gilbertiodendron brachystegioides, G. deweveri, Monopetalanthus letestui, M. microphyllus, and Brachystegia kennedyi. Along with Afrostyrax kamerunensis, <strong>the</strong> family Olacaceae dom<strong>in</strong>ates <strong>the</strong> middle-story <strong>of</strong> lowland forest with Strombosia grandiflora, S. pustulata, S. scheffleri, and Strombosiopsis tetandra be<strong>in</strong>g particularly abundant. The Ebenaceae are also a common component <strong>of</strong> <strong>the</strong> middle-story, particularly <strong>the</strong> genus Diospyros represented by D. crassiflora, D. preussii, D. sanzam<strong>in</strong>ika, D. simulans, D. suaveolens, and D. zenkeri. The Celastraceae are also abundant with many species <strong>of</strong> Salacia present. O<strong>the</strong>r abundant species are Angylocalyx zenkeri, Annickia chlorantha, Anonidium mannii, Polyalthia suaveolens, Calpocalyx d<strong>in</strong>klagei, Corynan<strong>the</strong> pachyceras, Cyrtogonone argentea, Dacryodes igaganga, Dactyladenia mannii, Dialium bip<strong>in</strong>densis, Discoglypremna caloneura, Garc<strong>in</strong>ia mannii, Grewia coriacea, Homalium dolichophyllum, Hypnodaphnis zenkeri, Isolona hexaloba, Monodora tenuifolia, Paus<strong>in</strong>ystalia macroceras, Plagiostyles africana, Polayalthia suaveolens, Protomegabaria stapfiana, Scottelia mimfiensis, Sor<strong>in</strong>deia grandifolia, Tabernaemontana crassa, Tapura africana, Treculia obovoidea, Usteria gu<strong>in</strong>eensis, Xylopia staudtii, and <strong>the</strong> stilt-rooted Santiria trimera and Uapaca gu<strong>in</strong>eensis. <strong>Takamanda</strong>: <strong>the</strong> Biodiversity <strong>of</strong> an African Ra<strong>in</strong>forest # <strong>of</strong> species Shannon©s Index (H©) Evenness (E) Common small trees <strong>in</strong> <strong>the</strong> understory <strong>in</strong>clude Antidesma lac<strong>in</strong>iatum var. lac<strong>in</strong>iatum, Baphia leptobotrys, B. nitida, Dorstenia tub<strong>in</strong>ata, Lasian<strong>the</strong>ra africana, Maesobotrya dusenii, Mareya micrantha, Mareyopsis longifolia, Massularia acum<strong>in</strong>ata, R<strong>in</strong>orea dentate, and Rothmannia lujae. Common herbs <strong>in</strong> <strong>the</strong> closed-canopy lowland forest are Costus englerianus, Dracaena camerooniana, D. phrynoides, Dorstenia mannii, D. barteri var. multiradiata, Mapania amplivag<strong>in</strong>ata, Renealmia polypus, and Scadoxus c<strong>in</strong>nabar<strong>in</strong>us. The Commel<strong>in</strong>aceae is also well represented <strong>in</strong> <strong>the</strong> herbaceous layer, notably Aneilmia ben<strong>in</strong>ense, Palisota ambigua, P. barteri, P mannii, P. capitata, and Pollia condensata. Groups <strong>of</strong> <strong>the</strong> crimson rosettes <strong>of</strong> <strong>the</strong> parasitic Thonn<strong>in</strong>gia sangu<strong>in</strong>ea are <strong>of</strong>ten seen on <strong>the</strong> forest floor. In <strong>the</strong> lowland forest, additional vegetation sub-types are dist<strong>in</strong>guishable; that is, river<strong>in</strong>e forest and extensive areas <strong>of</strong> secondary forest. Two plots were established <strong>in</strong> <strong>the</strong> river<strong>in</strong>e forest, but only qualitative collections were conducted <strong>in</strong> <strong>the</strong> secondary forests. 4.2.1.1 River<strong>in</strong>e forest Fisher©s Index Mean dbh (cm) Total BA Lowland ridge forest 6 491 103 3.95 0.85 39.7 30.0 34.7 Lowland ridge forest 7 498 98 3.82 0.83 36.0 32.8 42.1 Lowland ra<strong>in</strong>forest 10 414 90 3.86 0.86 34.8 29.7 28.7 Lowland ra<strong>in</strong>forest 11 428 113 4.01 0.85 50.1 25.4 21.7 Lowland (river<strong>in</strong>e) ra<strong>in</strong>forest 12 426 93 3.81 0.84 36.1 30.9 32.0 Lowland (river<strong>in</strong>e) ra<strong>in</strong>forest 13 477 118 4.03 0.84 50.2 29.3 32.1 Lowland ra<strong>in</strong>forest 14 406 83 3.62 0.82 31.0 35.6 40.5 Lowland ra<strong>in</strong>forest 15 438 91 3.89 0.87 33.7 31.1 33.2 Mid-altitude 8 527 64 3.29 0.79 19.1 24.4 24.6 Montane/savanna 9 523 74 3.59 0.84 23.5 21.3 18.6 The extensive dra<strong>in</strong>age pattern <strong>of</strong> <strong>the</strong> <strong>Takamanda</strong> region has given rise to large areas <strong>of</strong> lowland forest heavily <strong>in</strong>fluenced by seasonal <strong>in</strong>undation and periodic flood<strong>in</strong>g. This forest formation is relatively extensive with<strong>in</strong> <strong>the</strong>
Vegetation Assessment Figure 2. Structure and composition <strong>of</strong> lowland forest habitat <strong>in</strong> <strong>Takamanda</strong> Forest Reserve, Cameroon, from biodiversity plots, show<strong>in</strong>g relative density and relative basal area by (a) species, and (b) family. (a) Afrostyrax kamerunensis Strombosiopsis tetrandra Strombosia grandifolia Treculia obovoidea Irv<strong>in</strong>gia gabonensis Kla<strong>in</strong>edoxa gabonensis Cylicodricus gabunensis Tabernaemontana crassa Duboscia macrocarpa (b) Alstonia congensis Olacaceae Euphorbiaceae Leg-Mimosoideae Huaceae Irv<strong>in</strong>giaceae Leg-Caesalp<strong>in</strong>ioideae Apocynaceae Annonaceae Leg-Papilionoideae Sterculiaceae Relative Density Relative BA 0 2 4 6 8 10 Relative Density Relative BA 0 2 4 6 8 10 12 14 25 SI/MAB Series #8, 2003