Fisheries in the Southern Border Zone of Takamanda - Impact ...
Fisheries in the Southern Border Zone of Takamanda - Impact ...
Fisheries in the Southern Border Zone of Takamanda - Impact ...
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Vegetation Assessment <strong>of</strong> <strong>Takamanda</strong> Forest Reserve, Cameroon<br />
Chapter 3<br />
Terry C.H. Sunderland, James A. Comiskey, Simon Besong, Hyac<strong>in</strong>th<br />
Mboh, John Fonwebon and Mercy Abwe Dione<br />
1 Introduction<br />
Aside from South Africa, Cameroon is <strong>the</strong> most<br />
biologically rich country known to date on <strong>the</strong> African<br />
cont<strong>in</strong>ent (IUCN 1986). It encompasses an <strong>in</strong>tricate<br />
mosaic <strong>of</strong> diverse habitats, with moist, tropical forest<br />
dom<strong>in</strong>at<strong>in</strong>g <strong>in</strong> <strong>the</strong> south and sou<strong>the</strong>ast and cover<strong>in</strong>g 54%<br />
<strong>of</strong> <strong>the</strong> country (WCMC 1994), montane forest and alp<strong>in</strong>e<br />
savannah <strong>in</strong> <strong>the</strong> highlands, and sub-sahelian savannah<br />
and near desert <strong>in</strong> <strong>the</strong> far north (Letouzey 1968 and 1985,<br />
White 1983). These diverse habitats harbor more than<br />
9,000 species <strong>of</strong> plants, 160 <strong>of</strong> which are endemic<br />
(WCMC 1994). The majority <strong>of</strong> <strong>the</strong> endemic taxa are<br />
concentrated around Mount Cameroon and o<strong>the</strong>r<br />
highland areas.<br />
The forests <strong>of</strong> <strong>the</strong> cross-border region between<br />
Cameroon and Nigeria are also extraord<strong>in</strong>arily diverse,<br />
with a high degree <strong>of</strong> endemism (Davis et al. 1994). This<br />
area, where forest types are heavily <strong>in</strong>fluenced by<br />
dra<strong>in</strong>age patterns and topographical features, is <strong>the</strong> last<br />
refuge <strong>in</strong> Cameroon for <strong>the</strong> Cross River gorilla (Gorilla<br />
gorilla deihli; Sunderland-Groves et al. this volume).<br />
Compared to o<strong>the</strong>r forests <strong>in</strong> <strong>the</strong> Gu<strong>in</strong>eo-Congolian<br />
region, Cross River border forests are floristically<br />
diverse, with forest formations <strong>in</strong> <strong>Takamanda</strong> Forest<br />
Reserve (TFR) represent<strong>in</strong>g much <strong>of</strong> this diversity.<br />
This paper presents <strong>the</strong> f<strong>in</strong>d<strong>in</strong>gs <strong>of</strong> a vegetation<br />
study conducted <strong>in</strong> TFR between September 2000 and<br />
July 2001. For a more complete description <strong>of</strong> <strong>the</strong><br />
Reserve, see Sunderland-Groves et al. (this volume).<br />
1.1 Vegetation Surveys<br />
Botanical expeditions have been conducted <strong>in</strong> <strong>the</strong> larger<br />
study area s<strong>in</strong>ce <strong>the</strong> 1920s when numerous collections<br />
were made <strong>in</strong> <strong>the</strong> Obudu Plateau <strong>of</strong> Nigeria, which forms<br />
<strong>the</strong> western extension <strong>of</strong> <strong>the</strong> Bamenda Highlands <strong>of</strong><br />
Cameroon (Hall and Medler 1975). Keay (1979)<br />
provides descriptions <strong>of</strong> habitats identified dur<strong>in</strong>g a<br />
botanical expedition conducted <strong>in</strong> 1948. On <strong>the</strong><br />
Cameroon side, attention has focused on <strong>the</strong><br />
mounta<strong>in</strong>ous region, especially Mount Cameroon (for<br />
example, Cable and Cheek 1998, Hall 1973, Richards<br />
1963a and b). To <strong>the</strong> west and border<strong>in</strong>g on Nigeria’s<br />
Cross River National Park, Korup National Park has<br />
been <strong>the</strong> sett<strong>in</strong>g for extensive botanical and ecological<br />
work, <strong>in</strong>clud<strong>in</strong>g quantitative assessments (Newbery and<br />
Gartlan 1996), phenology (Newbery et al. 1996), and<br />
studies <strong>of</strong> nutrients and mycorrhizae (Gartlan et al. 1986,<br />
Newbery et al. 1988, Newbery et al. 1997). A<br />
prelim<strong>in</strong>ary botanical expedition to <strong>Takamanda</strong> was<br />
conducted by Etuge (1998). Two 1-ha biodiversity plots<br />
were established <strong>in</strong> 1997, <strong>the</strong> first <strong>in</strong> Cross River<br />
National Park and <strong>the</strong> second <strong>in</strong> <strong>the</strong> Okwangwo Division<br />
(Comiskey et al. unpublished data).<br />
Recently, <strong>the</strong> Smithsonian Institution established a<br />
forest dynamics plot where all trees with a diameter<br />
greater than 1 cm were measured <strong>in</strong> a 50-ha area <strong>in</strong><br />
Korup National Park (Schuster et al. 1999, Thomas et al.<br />
2003, Songwe et al. <strong>in</strong> press). To <strong>the</strong> north <strong>of</strong> Korup <strong>in</strong><br />
<strong>the</strong> Ejagham Forest Reserve, two 1-hectare biodiversity<br />
plots were established <strong>in</strong> 1997 (Sunderland et al. 1997).<br />
These plots were remeasured <strong>in</strong> 2002 and provide <strong>the</strong><br />
first quantitative <strong>in</strong>formation on dynamics <strong>of</strong> forests <strong>in</strong><br />
<strong>the</strong> region (Comiskey and Sunderland unpublished data).<br />
1.2 <strong>Takamanda</strong> Vegetation Assessment<br />
Until recently, <strong>the</strong>re was a considerable lack <strong>of</strong> detailed<br />
knowledge about vegetative composition <strong>in</strong> <strong>the</strong><br />
<strong>Takamanda</strong> area. As part <strong>of</strong> <strong>the</strong> present assessment, a<br />
SI/MAB Series #8, 2003, Pages 19 to 53