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87<br />

juvenile was moving to escape from us. Juveniles usually didn't move when approached but<br />

we removed al! observations of disturbed or moving juveniles from analyses (n = 8<br />

observations). Our telem<strong>et</strong>ry effort was concentrated in the daytime (90% of locations were<br />

obtained b<strong>et</strong>ween 9 am and 5 pm). We paid special attention to distribute sampling along the<br />

diurnal cycle evenly across indivi<strong>du</strong>als.<br />

Study <strong>de</strong>sign<br />

Each time we located an animal, we characterized the habitat where the animal was<br />

hiding and at a paired random location. For both the animal and the random location, we<br />

ma<strong>de</strong> one s<strong>et</strong> of mea<strong>sur</strong>es at the microhabitat scale (within 1 m of the animal or random<br />

location) and one at the local scale (within 15 m of the animal or random location). We first<br />

d<strong>et</strong>ennined wh<strong>et</strong>her selection for coyer occurred at one or both scales to verify our prediction<br />

1. Because the number of observations was low for some animaIs (mean ± SE: 21 ± 4<br />

observations, range: 2- 47, Table 5.3), we could not d<strong>et</strong>ermine habitat selection for each<br />

indivi<strong>du</strong>al. Rather, we d<strong>et</strong>ermined habitat selection for the popu<strong>la</strong>tion, taking into account the<br />

pseudo-replication in our analyses (following Fortin <strong>et</strong> al. 2005, see statistical analyses). To<br />

verify our prediction 2, we tested wh<strong>et</strong>her the use of coyer <strong>de</strong>pen<strong>de</strong>d on m<strong>et</strong>eorological<br />

conditions using data for the popu<strong>la</strong>tion and again taking pseudo-replication into account (see<br />

statistical analyses). Finally, we used mean coyer used by indivi<strong>du</strong>al juveniles (at both<br />

scales), sex, body mass on 10 June, mean daily movements, and mean distance to the mother<br />

as covariates in <strong>sur</strong>vival analyses in or<strong>de</strong>r to verify our prediction 3 and test for the effects of<br />

sex, body mass, mobility, and distance to the mother on <strong>sur</strong>vival time. Below we d<strong>et</strong>ail how<br />

we implemented this study <strong>de</strong>sign.<br />

Habitat selection<br />

When we located a juvenile, we first noted wh<strong>et</strong>her it was in a <strong>de</strong>n (usually rock <strong>de</strong>ns),<br />

on the ground, or in a tree (at least 30 cm above ground). ]f the juvenile was in a tree, we<br />

recor<strong>de</strong>d the species of the tree and mea<strong>sur</strong>ed its circumference at breast height. Then, we<br />

visually estimated «5%, 5-25%, 25-50%, 50-75%, >75%) the protective coyer (veg<strong>et</strong>ation or<br />

rock, Mysterud & Ostbye 1999) present within 1 m of the porcupine. Finally, we mea<strong>sur</strong>ed<br />

air temperature and wind speed at the animal location using a handheld thermom<strong>et</strong>erwindm<strong>et</strong>er<br />

(TF A 42.6000.06, TF A Dostmann Ltd Co., Germany). At the local scale (15 m-<br />

radius), we visually estimated «5%, 5-25%, 25-50%, 50-75%, >75%) tree, shrub (any

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