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59<br />

Third, we evaluated the behavioural response of porcupines to variations In Tc. We<br />

recor<strong>de</strong>d time spent outsi<strong>de</strong> of the <strong>de</strong>n, number of activity bouts, and a nocturnality in<strong>de</strong>x<br />

(equation 2 below) for each day (from 00:00 am to 24:00 pm) and compared them to the<br />

mean Tc of the day (recalcu<strong>la</strong>ted from continuous mea<strong>sur</strong>es of Ta, wind speed and n<strong>et</strong><br />

radiation ma<strong>de</strong> at our weather station, equation 3 below). We also compared the <strong>du</strong>ration of<br />

activity bouts to the mean Tc <strong>du</strong>ring the activity bout. Finally, we d<strong>et</strong>ermined which<br />

microhabitats were used when porcupines were outsi<strong>de</strong> of their <strong>de</strong>n, according to Tc<br />

mea<strong>sur</strong>ed in the stand at the time of observation. Because porcupines are sexually dimorphic,<br />

we tested for sexual differences in behaviour.<br />

M<strong>et</strong>eorological conditions<br />

Our automated weather station mea<strong>sur</strong>ed mr temperature In the sha<strong>de</strong> (Ta) using a<br />

temperature sensor (8-bit Temperature Smart Sensor, Hobo, Ons<strong>et</strong>) p<strong>la</strong>ced 1 m above ground,<br />

wind speed using a wind and direction sensor (Wind speed/Direction Smart Sensor, Hobo)<br />

p<strong>la</strong>ced 2 m above ground, and n<strong>et</strong> radiation (0-100 !lm) using a n<strong>et</strong> radiom<strong>et</strong>er sensor (mo<strong>de</strong>l<br />

NR-Lite N<strong>et</strong> Radiom<strong>et</strong>er, Kipp & Zonen) p<strong>la</strong>ced 1.5 m above ground. Ail sensors were<br />

connected to a datalogger (four channels Micro-station, Hobo) that collected data every<br />

minute.<br />

We p<strong>la</strong>ced temperature loggers (SmartButton Temperature Loggers, ACR systems lnc.)<br />

insi<strong>de</strong> nine <strong>de</strong>ns that were regu<strong>la</strong>rly used by porcupines <strong>du</strong>ring our long-term study, but that<br />

were not occupied <strong>du</strong>ring the sampling period. Loggers were p<strong>la</strong>ced on the ground as far as<br />

possible from the entrance of the <strong>de</strong>n (0.5 to 6.5 m <strong>de</strong>ep, <strong>de</strong>pending on <strong>de</strong>n structure).<br />

Loggers recor<strong>de</strong>d temperature every 15 minutes.<br />

Thermal mapping<br />

We worked <strong>du</strong>ring two winters to establish the thermal map: 29 January to 30 March<br />

2004 and 13 December 2004 to 8 February 2005. Following Bakken <strong>et</strong> al.(1983), we built<br />

two heated taxi<strong>de</strong>nnic mounts in the fonn of North American porcupines. To coyer the<br />

copper core we used pelts from animaIs that died naturally in our study area <strong>du</strong>ring winter<br />

2003. We built the mounts in a standing posture with the head facing down. We lined the<br />

insi<strong>de</strong> <strong>sur</strong>face of the hollow copper mould with 24 gauge Teflon-coated iron wire that was<br />

connected to an external control circuit and to a Campbell CR-21X data logger (Campbell<br />

Scientific). Electrical power was provi<strong>de</strong>d by four 6 V <strong>de</strong>ep discharge batteries in series.

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