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44<br />

Survival was a function of age and season, with time-<strong>de</strong>pen<strong>de</strong>nt w inter <strong>sur</strong>vival (Table<br />

3.4). Juveniles exhibited lower <strong>sur</strong>vival than indivi<strong>du</strong>als 2: 1-yr-old, whatever the season<br />

consi<strong>de</strong>red. Summer <strong>sur</strong>vival was hi gh and constant over the study period (mean monthly<br />

<strong>sur</strong>vival ± SE: 0.86 ± 0.06 for juveniles and 0.97 ± 0.01 for indivi<strong>du</strong>als 2: 1-yr-old) while<br />

winter <strong>sur</strong>vival was lower and variable from year to year (mean monthly <strong>sur</strong>vival: 0.53-0.83<br />

fOl·j uveniies and 0.84-0.96 for indivi<strong>du</strong>als 2:1-yr-old, Fig. 3.2). Winter precipitation<br />

eXp <strong>la</strong>ined a <strong>la</strong>rge pa11 (r 2 = 0. 87, Tabl e 3.5) of the yearly variations in winter <strong>sur</strong>vival (i.e. to<br />

inclu<strong>de</strong> this covariate <strong>de</strong>creased Alec by 5 units, Table 3.5). Snowfall al one eXp<strong>la</strong>ined a<br />

mo<strong>de</strong>rate amount of variation in winter <strong>sur</strong>vival whereas spring temperature exp<strong>la</strong>ined little<br />

variati on (Table 3.5). Winter <strong>sur</strong>vival was inversely re<strong>la</strong>ted to winter precipitation (Fig. 3.2)<br />

and, because of the in verse re<strong>la</strong>tionship b<strong>et</strong>ween winter precipitati on and so<strong>la</strong>r ilTadiance,<br />

positively re<strong>la</strong>ted to so<strong>la</strong>r irradiance (Fig. 3.2). We found no evi<strong>de</strong>nce ofa <strong>la</strong>gged effect (one<br />

and two years) of winter precipitation, snowfall, or spring temperature on winter <strong>sur</strong>vival<br />

(Table 3.6).<br />

Table 3.4 : Tests of the effects of sex, age (a, consi<strong>de</strong>ring two age c<strong>la</strong>sses: juveni les and 2: 1-<br />

yr-old), season (s) and year (y) on seasonal <strong>sur</strong>vival rates (~) in a popul ation of North<br />

American porcupines, Parc National <strong>du</strong> Bic, Québec, May 2000-May 2005. We Llsed data on<br />

159 indivi<strong>du</strong>als and mo<strong>de</strong>led sighting probabilities following mo<strong>de</strong>l selected in Table 3.3.<br />

Mo<strong>de</strong>l in bold represents the selected mo<strong>de</strong>l<br />

Biological meaning Notationt Deviance np* L'.A ICc**<br />

Constant <strong>sur</strong>vival

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