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36<br />

Hornfeldt 1994, Gese <strong>et</strong> al. 1996a, Se<strong>la</strong>s & Vik 2006). Prey living above the snow <strong>sur</strong>face<br />

usually suffer higher predation risk as snow <strong>de</strong>epens. We therefore expected winter predation<br />

rates on porcupines to increase in years of high snowfall/winter precipitation.<br />

3.3. Material and m<strong>et</strong>hods<br />

Study popu<strong>la</strong>tion<br />

We worked from May 2000 to May 2006 in a ca. 2 km 2 area of Parc National <strong>du</strong> Bic<br />

(48°20'N, 68°46'W, elevation 0-150m), Québec, Canada. The study area is fragmented by<br />

abandoned and cultivated fields and characterized by a rugged topography, abundance of<br />

natural rock <strong>de</strong>ns, and a mixed boreal forest. We captured porcupines in open fields <strong>du</strong>ring<br />

intensive capture sessions in May and August every year from 2000 to 2004 and in May only<br />

in 2005 and 2006 (Table 3.1). Each night of<strong>sur</strong>vey involved one to six observers (usually<br />

two) who patrolled the study area on foot or bicycle for one to nine hours (usually five). We<br />

sexed, weighed, aged (as juvenile, suba<strong>du</strong>lt, or a<strong>du</strong>lt using body mass in May, Berteaux <strong>et</strong> al.<br />

2005), and permanently marked porcupines upon capture with m<strong>et</strong>al ear tags (Be11eaux <strong>et</strong> al.<br />

•<br />

2005, Morin <strong>et</strong> al. 2005). We also used telem<strong>et</strong>ry to monitor indivi<strong>du</strong>als and fitted radio<br />

col<strong>la</strong>rs (Lotek SMRC-5RB VHF transmitters, Lotek Wireless Inc. , Newmark<strong>et</strong>, ON, Canada,<br />

L3Y 7B5) on 97 indivi<strong>du</strong>als for 23576 porcupine-days (Table 3.1).<br />

Parturition period is mid-May to <strong>la</strong>te June so we consi<strong>de</strong>red a popu<strong>la</strong>tion-year to start in<br />

May (e.g. year 2000 extends from May 2000 to April 2001). We estimated minimum<br />

popu<strong>la</strong>tion size for a given year as the number of different indivi<strong>du</strong>als observed b<strong>et</strong>ween May<br />

and April of the following year. Open fields are very attractive to porcupines for feeding in<br />

summer and porcupines were easily observed when using fields. We are therefore confi<strong>de</strong>nt<br />

that vi11ually ail porcupines present in the study area were captured or observed at least once<br />

each year. As a confirmation, we rarely found marked animais not observed one year and<br />

<strong>la</strong>ter relocated (only three and four animais, respectively in 2001 and 2002, which we ad<strong>de</strong>d<br />

to the popu<strong>la</strong>tion size for the year that they had not been observed).

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