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101<br />

However, we think protection from wind was not the main factor driving selection of covered<br />

microhabitats by juveniles because selection for coyer did not vary according to wind<br />

conditions (see below).<br />

Effects ofm<strong>et</strong>eorological conditions on the use of cover<br />

ln summer, use of co ver by animais usually results in a lower access to incoming<br />

radiation and thus to expo<strong>sur</strong>e to lower temperatures (Cook <strong>et</strong> al. 1998). Our results<br />

confirmed this pattern since air temperature was negatively corre<strong>la</strong>ted with protective cover<br />

at a given site. As predation was expected to drive habitat selection by juvenile porcupines,<br />

we predictedjuvenile porcupines would use open environments only un<strong>de</strong>r cold conditions<br />

(prediction 2). We found that animais increased their use of <strong>de</strong>ns when temperature <strong>de</strong>creased<br />

and when rain intensity increased. It was not <strong>sur</strong>prising that juveniles used their <strong>de</strong>n when<br />

rain intensity increased because a w<strong>et</strong> coat provi<strong>de</strong>s a poor insu<strong>la</strong>tion against cold<br />

temperatures (Kreith & B<strong>la</strong>ck 1980) so porcupines used <strong>de</strong>ns to avoid w<strong>et</strong>ting their coat.<br />

However, the fact that juveniles used <strong>de</strong>ns un<strong>de</strong>r co Id tempe rature conditions does not<br />

support our prediction that juveniles should use open areas on co Id days to maximize<br />

radiative heat gain. We trust juveniles were able to warm up their <strong>de</strong>ns when using them so<br />

that <strong>de</strong>ns constituted unique covered microhabitats that provi<strong>de</strong>d shelter against co Id<br />

temperatures, rain, and predators. Juveniles therefore did not face a tra<strong>de</strong>-off b<strong>et</strong>ween use of<br />

coyer and thennal expo<strong>sur</strong>e, but rather faced a tra<strong>de</strong>-off b<strong>et</strong>ween use of <strong>de</strong>n and non-<strong>de</strong>n<br />

microhabitats. Dens provi<strong>de</strong>d hiding coyer and protection against harsh weather conditions<br />

but no foraging Oppo11unities. As energy needs of juveniles increased (i.e. as their body mass<br />

increased with Julian date), porcupines <strong>de</strong>creased their use of <strong>de</strong>ns, possibly to increase the<br />

time spent foraging. An alternate (non-exclusive) hypothesis would be that juveniles<br />

<strong>de</strong>creased use oftheir <strong>de</strong>ns because the thermal environment became more favourable or<br />

because animais were able to thennoregu<strong>la</strong>te b<strong>et</strong>ter as the season advanced.<br />

Wind also <strong>influence</strong>d the use of <strong>de</strong>ns with juveniles being increasingly outsi<strong>de</strong> of their<br />

<strong>de</strong>n as wind speed increased. Biting insects are less abundant insi<strong>de</strong> than outsi<strong>de</strong> <strong>de</strong>ns<br />

(Comtois & Berteaux 2005) so that juveniles may weil use <strong>de</strong>ns on calm (i.e. non windy)<br />

days to avoid biting flies when insect harassment is high (i .e. when there is no wind). Outsi<strong>de</strong><br />

of the <strong>de</strong>n, we found no effect of air temperature on the use of coyer (at the microhabitat<br />

scale). However, juveniles used open microhabitats more often with increasing past rain

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