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28<br />

2.5. Discussion<br />

Searching<br />

Finding a<strong>du</strong>lt porcupines was re<strong>la</strong>tively easy in our study area in spring and at the end of<br />

autumn, when porcupines used open fields to forage. Juveniles were much more challenging<br />

to find . Had we limited to three the number of searches around each mother on a given year,<br />

and to 30 minutes the <strong>du</strong>ration of each search, we would have re<strong>du</strong>ced the total time spent<br />

sem·ching to 48.1 hours and the number of juveniles found per 100 hours of search wou Id<br />

have increased to 33 (16 juveniJes found in 48.1 hours). Hale and Fuller (1996) reported 10<br />

juveniles found after 176 one-hour searches around 23 a<strong>du</strong>lt females. Their overall success<br />

rate of 43.5 juveniles per 100 mothers is close to ours. Their searching m<strong>et</strong>hod and the<br />

number of sem"ches performed before finding each new juvenile was, however, not reported.<br />

Doing a maximum of three 30-minute searches per mother would increase search efficiency<br />

when many potential mothers are avai<strong>la</strong>ble. A greater search effort wou Id still be useful if<br />

few a<strong>du</strong>lt females are known and the number offound juveniles must be maximized. We<br />

recommend investing most of the effort searching the first 15 m around the mother,<br />

especially in a <strong>de</strong>nse forest habitat where searching a <strong>la</strong>rger radius is not efficient.<br />

Ascel1aining the repro<strong>du</strong>ctive status of females around which the searches are ma<strong>de</strong><br />

might increase success rate ofsearches. We consi<strong>de</strong>red a female to be <strong>la</strong>ctating ifsome liquid<br />

COLI Id be manually extracted from nipples. This was not al ways reliable. For example, we<br />

ma<strong>de</strong> Il lInsuccessful searches around a female who <strong>la</strong>ter proved to be pregnant <strong>du</strong>ring these<br />

searches (the twelfth search yiel<strong>de</strong>d a newborn porcupine; see Fig. 2.3a). An alternative,<br />

probably b<strong>et</strong>ter m<strong>et</strong>hod to d<strong>et</strong>ermine repro<strong>du</strong>ctive status consists in checking nipples after<br />

stimu<strong>la</strong>tion of milk release by injection of oxytocin (Roze 1989, Hale & Fuller 1996). We<br />

tried this m<strong>et</strong>hod in the first stages of our study but abandoned it after a few unsuccessful<br />

tests. We, however, suspect that our <strong>la</strong>ck of experience with porcupine handling biased<br />

negatively this early test.<br />

Capu/ring<br />

The most efficient capture m<strong>et</strong>hod usually was the dip n<strong>et</strong>. Use of a dip n<strong>et</strong> had already<br />

been repol1ed (Zimmerling & Croft 2001 , Zimmerling 2005) but authors did not <strong>de</strong>scribe<br />

equipment nor did they evaluate m<strong>et</strong>hod efficiency. A single experienced observer can safely

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