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19<br />

information is <strong>la</strong>cking in the published literature, and we finally suggest four research<br />

obj ecti ves that will structure the rest of the paper.<br />

Porcupines usually live in forests where they are difficult to locate. Hale and Fuller<br />

( 1999) report low success wh en walking along transects to search for porcupines. Sweitzer<br />

and Berger (1997) worked in re<strong>la</strong>tively open habitats and were successful at locating<br />

porcupines using trained dogs and night-vision equipment. However, by far the most efficient<br />

<strong>de</strong>scribed technique to find porcupines involves snow tracking to locate <strong>de</strong>ns where<br />

porcupines concentrate their activity in winter (Strick<strong>la</strong>n <strong>et</strong> al. 1995, Zimmerling 2005).<br />

Capture techniques inclu<strong>de</strong> constraining porcupines in a garbage can (Craig & Keller<br />

1986), climbing trees to catch indivi<strong>du</strong>als using heavy gloves (Roze 1989, Strick<strong>la</strong>n <strong>et</strong> al.<br />

1995), p<strong>la</strong>cing cage-type live traps outsi<strong>de</strong> occupied <strong>de</strong>ns, trees or on well-used paths (Hale<br />

& Fuller 1996, Griesemer <strong>et</strong> al. 1999, lise & Hellgren 2001, Zimmerling & Craft 2001),<br />

n<strong>et</strong>ting the animal before (Zimmerling & Croft 2001 , Zimmerling 2005) or after (lIse &<br />

Hellgren 2001) chemical immobilization, and using heavy gloves after immobilizing animaIs<br />

in trees or <strong>de</strong>ns using a jabstick (Sweitzer & Berger 1992, Griesemer <strong>et</strong> al. 1999, lise &<br />

Hellgren 2001) or a blow dart (Griesemer <strong>et</strong> al. 1999).<br />

Marking techniques involve m<strong>et</strong>al ear tags (Craig & Keller 1986, Zimmerling & Croft<br />

2001) to which are som<strong>et</strong>imes attached colored p<strong>la</strong>stic tags to allow i<strong>de</strong>ntification at a<br />

distance (Sweitzer & Berger 1992, Griesemer <strong>et</strong> al. 1999), spraying tail with enamel paint<br />

(Sweitzer & Berger 1992), tattooing (Griesemer <strong>et</strong> al. 1999), fitting radio transmitter using<br />

hm·ness and back pack (Craig & Keller 1986), or radiocol<strong>la</strong>ring (e.g. Roze 1987, Sweitzer &<br />

Berger 1992, Griesemer <strong>et</strong> al. 1999, lIse & Hellgren 2001). F<strong>la</strong>gs have been attached to<br />

antennas of radio col<strong>la</strong>rs to ease i<strong>de</strong>ntification of indivi<strong>du</strong>als at a distance (Griesemer <strong>et</strong> al.<br />

1999).<br />

Published techniques to search for, capture, and mark porcupines represent a fraction of<br />

those that have been tried by biologists, and m<strong>et</strong>hod sections of publications usually do not<br />

quantify efficiency of techniques. Griesemer <strong>et</strong> al. (1999) evaluate some m<strong>et</strong>hods to capture<br />

and mark a<strong>du</strong>lt porcllpines. They do not <strong>de</strong>al with jllveniles, however, and some of their<br />

reported m<strong>et</strong>hods nee<strong>de</strong>d improvement. Specifically, the m<strong>et</strong>hod they used for mm"king did<br />

not allow i<strong>de</strong>ntification at a distance and was not long-<strong>la</strong>sting (25% of indivi<strong>du</strong>als had lost ail<br />

tags 6-18 months after tagging). AIso, 7% of their radio col<strong>la</strong>rs created wOllnds (Griesemer <strong>et</strong>

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