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70 -..c -~20 .- nl '"C nl c: ~ 15 . Ū) - -~ o ai 10 a. U) Q) E -c: nl Q) ~ 5 o +-1- - 2, 2 7,44 7,33 0 ~ 80 1 - 60 - 40 - 20 o ~ ~ o > nl ..c Q) .c 0) c: '"C Q) Q) -o Q) o c: Q) ~ ~ ~ o o s:::: nl Q) ~ Operative tel1lJerature (OC) Figure 4.3 : Time spent outside of a den (mean ± SE) in a day for male (open circles) and female (fi lied circles) North Ameri can porcupines and OCCU1Tence offeeding behaviour (mean ± SE) when in a coniferous tree (fi lled squares: males and females pooled since difference between sexes was not significant) as a function of operative temperature, Parc National du Bic, Québec, January to April, 2004-2005. Sample sizes are specified near each symbol as: number of individuals, number of observations on those individuals.
71 Number and duration of activity bouts Mean number of activity bouts was lAO ± 0.07 (range: 1.07 to 1.91 , n = 14 individuals and 679 observations) and the me an duration of activity bouts was 5.0 ± OA h (range: 2.8 to 7.3 h, n = 14 individuals and 966 observations). Number ofactivity bouts increased wh en Tc decreased (F(I,664) = 20.59, p
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UNIVERSITÉ DU QUÉBEC À RIMOUSKI
Page 3 and 4:
Remerciements Cette thèse n'aurait
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Table des matières LISTE DES TABLE
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Liste des tableaux Chapitre 3 Table
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VB Table 4.2 Power consumed (in Wat
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Liste des figures Chapitre 1 Figure
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XI Figure 2.3 Number oftimes ajuven
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Xlll Figure 4.3 Time spent outside
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Liste des annexes Annexe 1 Descript
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Chapitre 1 Introduction 1.1 Impact
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3 L ' importance relative des facte
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5 des populations ne donne qu' une
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7 (Tympanuchus pallidicinctus) les
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9 1940 1000 1960 139---------- c ~
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1 1 (Roze 1987). Ils perdent 10 à
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13 140 a) 1 20 - o -l- 2000 2001 20
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15 manipulations et la pose d'émet
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Chapitre 2 Handling North American
Page 37 and 38: 19 information is lacking in the pu
Page 39 and 40: 21 We searched for juvenile porcupi
Page 41 and 42: 23 Switzerland). To check sex and r
Page 43 and 44: 25 transmitters fo r larger units s
Page 45 and 46: 27 8 a) 7 "0 c: :l 0 6 -If) ~ c: 5
Page 47 and 48: 29 record sex, reproducti ve status
Page 49 and 50: Chapitre 3 Predation as a possible
Page 51 and 52: 33 3.2. Introduction A growing numb
Page 53 and 54: 35 predation risk. To do so, we tes
Page 55 and 56: 37 Table 3.1 : Monitoring effort ac
Page 57 and 58: 39 winter precipitation (total prec
Page 59 and 60: 41 emaciated), road kills, dead fro
Page 61 and 62: 43 Seasonal survival rates The sele
Page 63 and 64: 45 >. 1.0 0.9 ..... 0.8 .c ta .c 0.
Page 65 and 66: 47 Causes of mortality We assigned
Page 67 and 68: 49 100 - 90 a) 80 1/) c: .2 iii ...
Page 69 and 70: 51 too simplistic to detect complex
Page 71 and 72: 53 correlated with winter survival
Page 73 and 74: 55 4.1. A bstract Because the clima
Page 75 and 76: 57 Ta, first because of the effects
Page 77 and 78: 59 Third, we evaluated the behaviou
Page 79 and 80: 61 pattern of den use over one or t
Page 81 and 82: 63 two modali ties, Table 4.1, mode
Page 83 and 84: 65 feeding behaviour for the popula
Page 85 and 86: 67 of Tc we considered. However, we
Page 87: 69 Calculating TefrOIn Ta, wind spe
Page 91 and 92: 73 8 o _ --L-_ 0 Operative temperat
Page 93 and 94: 75 Beschta 2004). Predation rates w
Page 95 and 96: 77 in our population (chap. 3). Mal
Page 97 and 98: 79 Tableau 4.5 : Pourcentage d'obse
Page 99 and 100: 81 associées avec le temps de surv
Page 101 and 102: 83 5.1. Abstract ln many mammals, j
Page 103 and 104: 85 use of coyer influenced summer s
Page 105 and 106: 87 juvenile was moving to escape fr
Page 107 and 108: 89 individual (using approximations
Page 109 and 110: 91 den use (treated as a binary var
Page 111 and 112: 1 93 20 - 18 ~ a) DAvailable .Used
Page 113 and 114: 95 80 -, 70 l DAvailable .Used 60 -
Page 115 and 116: 97 Table 5.3: Use of coyer by juven
Page 117 and 118: 99 juveniles. At the microhabitat s
Page 119 and 120: 101 However, we think protection fr
Page 121 and 122: 103 we found that most mortalities
Page 123 and 124: Chapitre 6 Conclusion Dans le cadre
Page 125 and 126: 107 porcs-épics fréquentent les c
Page 127 and 128: 109 épies, via un impact des condi
Page 129 and 130: II I Hiver E n hi ver, on montre qu
Page 131 and 132: 113 compromis auquel les animaux fo
Page 133 and 134: 11 5 En été, la mortalité des je
Page 135 and 136: 117 pékan dans la région. On mont
Page 137 and 138: 119 Annexe 2 Étalonnage des modèl
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121 Références Aanes, R. , & R. A
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Bult, A., & C. B. Lynch. 1997. Nest
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125 Creel, S. , 1. Winnie, B . Maxw
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127 Fortin, D., & G. Gauthier. 2000
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129 Hik, D. S. 1995. Does risk of p
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Lent, P. C. 1974. Mother-infant rel
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133 Murray, D. L. , & S. Boutin. 19
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au Québec depuis 1984. Ministère
Page 155 and 156:
137 Proceedings of the Royal Societ
Page 157 and 158:
139 Tsiropoula, G. 2003. Signatures
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