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Literature review: Impact of Chilean needle grass ... - Weeds Australia

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on basalt duplex soils had the highest probablity <strong>of</strong> destruction but also supported the largest number <strong>of</strong> <strong>grass</strong>lands. These are<br />

more readily cultivated than stony and uniform soils and are relatively high in nutrients. In terms <strong>of</strong> the road class <strong>of</strong> the nearest<br />

roads, <strong>grass</strong>lands on local roads had the highest probability <strong>of</strong> destruction, followed by patches on highways, possibly because<br />

local roads are more likely to be grazed, illegally cropped or sprayed with herbicides. In terms <strong>of</strong> land tenure, <strong>grass</strong>lands on<br />

Crown land were most likely to have been destroyed (mostly by weed invasion), those on private land were less likely to have<br />

been destroyed than those on roadsides, and no railway line <strong>grass</strong>lands were lost. However the number <strong>of</strong> private <strong>grass</strong>lands in<br />

1984 was severely underestimated, and it is likely that in reality large numbers were probably destroyed during the period.<br />

Country Fire Authority brigades also had a major impact on destruction: those brigades area where management <strong>of</strong> <strong>grass</strong>lands<br />

had changed from burning to managment using herbicides experienced the highest levels <strong>of</strong> destruction.<br />

McDougall (1987) argued that management requirements for remnants needed to be site specific, depending on the particular<br />

weed problems and specific conservation requirements, and that much remained to be learned. He acknowledged that the effects<br />

<strong>of</strong> fire on the inter-tussock herbs was then largely unknown, but recommended burning at 3-5 year intervals between early<br />

October and mid-December to reduce weeds and prevent overdominance by T. triandra, except at sites with spring-flowering<br />

rare or endangered species, for which late summer or autumn fires were appropriate. The main management issues for these<br />

<strong>grass</strong>lands currently include inadequate knowledge <strong>of</strong> threatened taxa, management <strong>of</strong> introduced <strong>grass</strong>es, prevention <strong>of</strong> new<br />

weed incursions and generalised ecosystem degradation (Groves and Whalley 2002).<br />

Loss <strong>of</strong> unreserved remnants continues through development for housing and agriculture. Degradation <strong>of</strong> ‘protected’ remnants<br />

also continues, including decline in quality, loss <strong>of</strong> species etc., as a result <strong>of</strong> invasive species, pollution at the local and global<br />

levels (e.g. N and CO 2 enrichment <strong>of</strong> the atmosphere), alteration <strong>of</strong> hydrological processes, other changed ecological processes,<br />

and inappropriate or inadequate management.<br />

South <strong>Australia</strong><br />

South <strong>Australia</strong> once had immense areas <strong>of</strong> temperate native <strong>grass</strong>land and <strong>grass</strong>y woodland in what is now the agricultural zone<br />

(Davies 1997). The <strong>grass</strong>y vegetation in that State occurred mainly on plateaus and in broad valleys on lower slopes from the<br />

Orroroo and Peterborough areas in the southern and western Flinders Ranges through the Mount L<strong>of</strong>ty Ranges to Peterborough<br />

and Murray Bridge. Smaller areas occurred on the sub-coastal plain <strong>of</strong> the South East between Bordertown and the Victorian<br />

border, on the southern Eyre Peninsula, the Yorke Peninsula, and on basalt soils near Mount Gambier (Lunt et al. 1998).<br />

However their composition and previous distribution is unclear (Davies 1997). As in Victoria, recognition that <strong>grass</strong>lands existed<br />

in some areas has been slow in coming. Lange (1983) failed to acknowledge the existence <strong>of</strong> <strong>grass</strong>lands in the South East. The<br />

<strong>grass</strong>es there, he noted, “do not form savannah; usually they grow sparsely” (p. 99). In the mid 1990s half <strong>of</strong> the significant<br />

South <strong>Australia</strong>n remnants were on private land, one third on roadsides and one sixth on Crown land (Kirkpatrick et al. 1995).<br />

Exotic plants have been reported to comprise 30% <strong>of</strong> the South <strong>Australia</strong>n temperate <strong>grass</strong>land vascular flora (Kirkpatrick et al.<br />

1995).<br />

Moore (1993) mapped these <strong>grass</strong>lands as Temperate Short<strong>grass</strong> (Austrodanthonia-Austrostipa-Enneapogon) formations,<br />

extending from south <strong>of</strong> Adelaide through the Mount L<strong>of</strong>ty Ranges to north <strong>of</strong> Port Augusta, and in parts <strong>of</strong> the South East.<br />

Two extant temperate <strong>grass</strong>land communities are under threat or inadequately conserved: Lomandra effusa/Lomandra multiflora<br />

subsp. dura (open) tussock <strong>grass</strong>land (co-dominant with Austrodanthonia and Austrostipa spp.), known as Mat-rush Grassland<br />

(Scented Mat-rush and Stiff Mat-rush Lomandra dura according to Lunt et al. 1998), or Iron<strong>grass</strong> communities (Carter et al.<br />

1993) and Austrodanthonia/Themeda tussock <strong>grass</strong>land. The former occurs on similar soil types and under similar ecological<br />

conditions to other temperate <strong>grass</strong>land communities in <strong>Australia</strong>, so despite the dominants not being <strong>grass</strong>es, the formations are<br />

considered in ecophysiological terms to be temperate <strong>grass</strong>lands (Carter et al. 1993). It is most common on skeletal soils in the<br />

eastern Mount L<strong>of</strong>ty Ranges and the Mid North (Kirkpatrick et al. 1995). Both communities were mainly found in the Mid<br />

North, in the Clare-Port Pirie-Peterborough region but have largely been destroyed by cropping and grazing. Grass dominated<br />

communities may have occurred near Adelaide and more widely in the lower South East, but are extremely poorly conserved<br />

(Lunt et al. 1998, Davies 1997).<br />

Areas <strong>of</strong> T. triandra <strong>grass</strong>land are “suspected to have existed” in the lower South East, “primarily on basalt soils (e.g. around<br />

Mount Schanck and Mount Gambier)”(Foulkes and Heard 2003 p. 48). These areas were described by early settlers as lighted<br />

wooded with Acacia melanxoylon R.Br, and eucalypts.<br />

A number <strong>of</strong> other <strong>grass</strong>land community types are present in South <strong>Australia</strong>, including Austrostipa <strong>grass</strong>lands and Murray<br />

Lakes <strong>grass</strong>lands (Davies 1997). The Austrodanthonia caespitosa – Enchylaena tomentosa Marsh Margin Grassland occurred on<br />

the margins <strong>of</strong> the Murray lakes, and “only survives in small roadside fragments” (Kirkpatrick et al. 1995). A Themeda-<br />

Geranium retrorsum-Arthropodium strictum community occurs in the Mount L<strong>of</strong>ty Ranges in the Mid North (Kirkpatrick et al.<br />

1995). Mathison (2004) stated that South <strong>Australia</strong>n native <strong>grass</strong>lands are not climax communities but transitional states and can<br />

radically change their nature when there is a change in the management regime. Stafford (1991) provided some descriptive<br />

information on secondary <strong>grass</strong>land (transitional to woodland) developed in the East Torrens district after cessation <strong>of</strong> cropping<br />

and grazing. This probably constituted ‘enriched <strong>grass</strong>land’ in the sense <strong>of</strong> McIntyre and Lavorel (2007).<br />

Approximately half <strong>of</strong> the 20 most abundant vascular plant species in South <strong>Australia</strong>n <strong>grass</strong>lands are exotic and annual <strong>grass</strong>es<br />

including Avena spp. Bromus spp. and Vulpia spp. are amongst the most invasive (Lenz et al. 2003)<br />

Tasmania<br />

As in other areas <strong>of</strong> southern <strong>Australia</strong>, the natural <strong>grass</strong>lands <strong>of</strong> Tasmania were rapidly occupied by European settlers (Benson<br />

and Redpath 1997), however they are better reserved than mainland <strong>grass</strong>lands (Kirkpatrick et al. 1995). Tasmanian non-alpine<br />

<strong>grass</strong>lands occur mostly on valley bottoms, on heavy clays or alluvial soils and lower slopes on shallow rocky soils, mostly in<br />

the drier Midlands” (Lunt et al. 1998). Moore (1993) mapped a narrow band <strong>of</strong> Temperate Short<strong>grass</strong> (Austrodanthonia-<br />

138

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