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climate change on UAE - Stockholm Environment Institute-US Center

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is based <strong>on</strong> a theoretical understanding of<br />

system dynamics (bottom-up) or if it is built <strong>on</strong><br />

empirical observati<strong>on</strong>s (top-down).<br />

Models built from the top down are more<br />

comm<strong>on</strong>, if <strong>on</strong>ly because the line between an<br />

experimental c<strong>on</strong>struct and a model is blurred<br />

when using empirical data. Empirical models can<br />

be as simple as deriving a functi<strong>on</strong> to describe<br />

a relati<strong>on</strong>ship through a set of variables, or<br />

can be as complex a system which predicts<br />

spatial patterns of vegetati<strong>on</strong> under changing<br />

<str<strong>on</strong>g>climate</str<strong>on</strong>g> c<strong>on</strong>diti<strong>on</strong>s (see the CASA model,<br />

below). Empirical models may be methods of<br />

interpreting or simplifying datasets with rich<br />

temporal or spatial informati<strong>on</strong>, such as l<strong>on</strong>g<br />

time series or satellite data; or may use dense<br />

datasets to compile statistical relati<strong>on</strong>ships,<br />

which may then be used for predictive purposes.<br />

The advantages of empirically-based models is<br />

that they can be relatively simple to c<strong>on</strong>struct<br />

and interpret, are often highly explicit in their<br />

assumpti<strong>on</strong>s, and, most importantly, are based<br />

directly <strong>on</strong> data.<br />

Broadly, the bottom-up modeling approach<br />

relies <strong>on</strong> established theories <strong>on</strong> how individual<br />

comp<strong>on</strong>ents of an ecosystem operate at the<br />

micro-scale. These mechanistic models are<br />

often built to be as general as possible, such<br />

that they are not c<strong>on</strong>strained by data (collected<br />

by fallible observers) or limited by the way<br />

communities and ecosystems are structured<br />

today. The point of these models is to explore<br />

relati<strong>on</strong>ships between ecosystem comp<strong>on</strong>ents<br />

and forcing factors, understand dynamics, and<br />

impose c<strong>on</strong>diti<strong>on</strong>s <strong>on</strong> a simulated ecosystem<br />

which may not exist today. Some of the most<br />

developed versi<strong>on</strong>s of these models are able<br />

to predict the structure and functi<strong>on</strong> of major<br />

biomes from first principles (i.e. photosynthesis,<br />

respirati<strong>on</strong>, and nutrient requirements), and are<br />

now being used to explore how <str<strong>on</strong>g>climate</str<strong>on</strong>g> <str<strong>on</strong>g>change</str<strong>on</strong>g>,<br />

land use, and disturbance may impact future<br />

biomes.<br />

Model limitati<strong>on</strong>s<br />

All ecosystem models are severely limited by<br />

scale, scope, and assumpti<strong>on</strong>s. Key aspects of<br />

each are briefly described below.<br />

Scale: The most fundamental processes in<br />

an ecosystem occur at micro-scales, where<br />

photosynthesis and respirati<strong>on</strong> occur,<br />

nutrients are utilized, and water is cycled. There<br />

are, however, also important processes which<br />

happen at the scale of the leaf (for example,<br />

growth, senescence, shading, herbivory), the<br />

stem (individual mortality, light and water<br />

availability), the patch (disturbances), the<br />

community (competiti<strong>on</strong>), and the regi<strong>on</strong><br />

(<str<strong>on</strong>g>climate</str<strong>on</strong>g>, light availability). The levels to<br />

which these processes are simulated are<br />

computati<strong>on</strong>ally and data limited, and many<br />

processes operate across scales.<br />

Scope: The broader a model strives to be,<br />

the more general its assumpti<strong>on</strong>s must<br />

become. To simulate a single type of biome<br />

effectively, <strong>on</strong>e might choose to model or<br />

simulate a modest number of individual<br />

floristic species with known characteristics;<br />

to then include yet more diverse biomes in<br />

the model, <strong>on</strong>e often has to reduce the level<br />

of detail down to functi<strong>on</strong>al plant types<br />

rather than individual species. Models which<br />

are global in scope often reduce plant types<br />

down to simple plant functi<strong>on</strong>al types which<br />

distinguish between physiognomy (tree or<br />

grass), leaf form (broadleaf or needle-leaf),<br />

leaf l<strong>on</strong>gevity (evergreen or deciduous), and<br />

photosynthetic pathway (C 3<br />

or C 4<br />

) (i.e. Wang<br />

et al., 2004). Models which effectively capture<br />

global-scale dynamics may be ineffective or<br />

irrelevant for studies at the sub-biome scale.<br />

Assumpti<strong>on</strong>s: Every form of ecosystem model<br />

has (or should have) a well developed list of<br />

general assumpti<strong>on</strong>s. Empirical models, for<br />

example, often assume that relati<strong>on</strong>ships<br />

between correlated variables are causal, or<br />

at least replicable, and rarely model discrete<br />

pathways or mechanisms. First-principles<br />

(bottom-up) models are rarely entirely<br />

mechanistic: at some level, even the most<br />

rigorous models simplify processes and<br />

mechanisms with empirical relati<strong>on</strong>ships.<br />

9.2 Types of ecosystem models<br />

Ecosystem models span a wide range of<br />

functi<strong>on</strong>s, but several types may be useful for<br />

evaluating the impacts of <str<strong>on</strong>g>climate</str<strong>on</strong>g> <str<strong>on</strong>g>change</str<strong>on</strong>g> <strong>on</strong><br />

ecosystems and biodiversity in the <strong>UAE</strong>.<br />

Impacts, Vulnerability & Adaptati<strong>on</strong> for<br />

Dryland Ecosystems in Abu Dhabi<br />

177

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