climate change on UAE - Stockholm Environment Institute-US Center
climate change on UAE - Stockholm Environment Institute-US Center
climate change on UAE - Stockholm Environment Institute-US Center
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First principles ecosystem<br />
models: potential vegetati<strong>on</strong> and<br />
disturbance<br />
Mechanistic models are designed to simulate<br />
the structure, functi<strong>on</strong>, and (sometimes)<br />
dynamics of ecosystems based <strong>on</strong> a “first<br />
principles” understanding of the comp<strong>on</strong>ents<br />
of the ecosystem. Exclusively, these models<br />
simulate and follow <str<strong>on</strong>g>change</str<strong>on</strong>g>s in vegetati<strong>on</strong><br />
compositi<strong>on</strong> <strong>on</strong>ly, and do not c<strong>on</strong>sider fauna,<br />
except occasi<strong>on</strong>ally as numerical agents of<br />
disturbance. The bottom-up approach is an<br />
effective way of exploring how a biome forms,<br />
and which climatic and competitive features<br />
drive the equilibrium compositi<strong>on</strong> of a biome.<br />
There are no standard structures for mechanistic<br />
models, but many do share comm<strong>on</strong> features.<br />
The mechanistic comp<strong>on</strong>ent is usually a<br />
set of equati<strong>on</strong>s describing carb<strong>on</strong> balance<br />
(photosynthesis and respirati<strong>on</strong>) with available<br />
light and water for a small variety of plant<br />
functi<strong>on</strong>al types (i.e. grasses, shrubs, trees,<br />
and deciduous and c<strong>on</strong>iferous species). Various<br />
levels of sophisticati<strong>on</strong> in these models may<br />
describe important ecosystem comp<strong>on</strong>ents,<br />
depending <strong>on</strong> the questi<strong>on</strong> at hand:<br />
Water availability and transfer: water<br />
infiltrati<strong>on</strong> into the soil and uptake by roots<br />
(CARLUC, Hirsch et al., 2004)<br />
Physical structures: tall trees deprive shorter<br />
shrubs and grasses of light (ED, Moorcroft et<br />
al., 2001)<br />
Disturbances: fires, windstorms, and other<br />
destructive events (IBIS, Foley et al., 1998;<br />
ED)<br />
Nutrient dynamics: available nitrogen in<br />
soils, roots, stems, and leaves (TEM, Tian et<br />
al., 1998)<br />
Seas<strong>on</strong>ality: <str<strong>on</strong>g>change</str<strong>on</strong>g>s in leaf density,<br />
senescence (CASA, Potter et al., 2004)<br />
These mechanistic models are useful for<br />
understanding how large scale <str<strong>on</strong>g>change</str<strong>on</strong>g>s in<br />
<str<strong>on</strong>g>climate</str<strong>on</strong>g> or other abiotic factors will <str<strong>on</strong>g>change</str<strong>on</strong>g><br />
biome locati<strong>on</strong>s or biomass, or tracing complex<br />
feedback mechanisms (such as how shifts in<br />
vegetati<strong>on</strong> abundance impact <str<strong>on</strong>g>climate</str<strong>on</strong>g> patterns,<br />
Wang et al., 2004). These models, however, are<br />
difficult to apply at small scales and lack the<br />
ability to discriminate <str<strong>on</strong>g>change</str<strong>on</strong>g>s in compositi<strong>on</strong><br />
more detailed than basic functi<strong>on</strong>al types. In<br />
additi<strong>on</strong>, mechanistic models can <strong>on</strong>ly describe<br />
a limited degree of complexity, and may neglect<br />
detailed, yet critical, interacti<strong>on</strong>s (such as<br />
nutrient or water flow between clustered<br />
shrubs and grasses in a semi-arid system, or<br />
different phenological resp<strong>on</strong>ses to seas<strong>on</strong>ality<br />
and drought).<br />
This class of model may be useful in determining<br />
<str<strong>on</strong>g>climate</str<strong>on</strong>g> impacts <strong>on</strong> the <strong>UAE</strong> if the predominant<br />
questi<strong>on</strong> is in regard to biomass, ecosystem<br />
feedback cycles, or how an ecosystem might be<br />
structured in the <strong>UAE</strong> without an anthropogenic<br />
influence.<br />
Bioclimatic envelope models<br />
Bioclimatic envelope models are designed to<br />
explore how species ranges may shift in resp<strong>on</strong>se<br />
to <str<strong>on</strong>g>climate</str<strong>on</strong>g> <str<strong>on</strong>g>change</str<strong>on</strong>g>. It has l<strong>on</strong>g been understood<br />
that <str<strong>on</strong>g>climate</str<strong>on</strong>g> (precipitati<strong>on</strong> and temperature)<br />
str<strong>on</strong>gly c<strong>on</strong>trols the ability of certain species<br />
and functi<strong>on</strong>al types to survive and thrive<br />
(Pears<strong>on</strong> and Daws<strong>on</strong>, 2003), and in fact, <strong>on</strong>e<br />
of the central tenants of biogeographical niche<br />
theory is that an ecological niche can be defined<br />
by the envir<strong>on</strong>mental variables which affect a<br />
species. Major biomes, and even biogeographical<br />
boundaries within these biomes, are largely<br />
defined climatically. Individual species may have<br />
a narrow range of acceptable <str<strong>on</strong>g>climate</str<strong>on</strong>g>s, or an<br />
envelope in which they are typically found (the<br />
“realized niche”) or should be found based <strong>on</strong><br />
known biological functi<strong>on</strong>s (the “fundamental<br />
niche”). Bioclimatic modeling asserts that we<br />
can anticipate the ecosystem impact of <str<strong>on</strong>g>climate</str<strong>on</strong>g><br />
<str<strong>on</strong>g>change</str<strong>on</strong>g> <strong>on</strong> species ranges (at the regi<strong>on</strong>al scale)<br />
by determining the new bounds <strong>on</strong> bioclimatic<br />
envelopes.<br />
Bio<str<strong>on</strong>g>climate</str<strong>on</strong>g> envelope modeling suffers from<br />
at least three shortcomings (see Pears<strong>on</strong> and<br />
Daws<strong>on</strong>, 2003):<br />
Competiti<strong>on</strong>: how a species might thrive in<br />
an envir<strong>on</strong>ment and how it actually interacts<br />
in its community can be very different; if a<br />
species is n<strong>on</strong>-competitive within its <str<strong>on</strong>g>climate</str<strong>on</strong>g><br />
envelope, it may not be found in the new<br />
envir<strong>on</strong>ment.<br />
Adaptati<strong>on</strong>: it may be more effective for a<br />
178<br />
Climate Change Impacts, Vulnerability & Adaptati<strong>on</strong>