Catalog of species-group names of Recent and fossil Scaphopoda ...

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Steiner G. & Kabat A. R.Formation. Marambio Island, James Ross GroupIslands, Antarctica. Holotype CIRGEO (Centro deInvestigaciones en Recursos Geológicos) PI 242(now housed in Museo Argentino de CienciasNaturales, Buenos Aires, fide G. Pastorino in litt.17.I.2001).wilsoni, Dentalium Fraas, 1867. Not a scaphopod. SeeOther taxa described as Scaphopoda.† wollumbillaensis, Dentalium Etheridge in Jack &Etheridge, 1892: 483. Cretaceous. Wollumbilla,Queensland, Australia. Neotype PIRS M2430 (designatedby Stilwell 1999: 220). Replacement namefor Dentalium lineatum Moore, 1870, a juniorhomonym of Dentalium lineatum Gueranger, 1853.Objective junior synonym: Dentalium mooreiPilsbry & Sharp, 1898. Fustiaria wollumbillaensis(Etheridge in Jack & Etheridge, 1892) (fideLudbrook 1966: 188; fide Stilwell 1999: 220).woodhousae, Cadulus Lamprell & Healy, 1998: 163,164, figs 167D, 168G, 172. Recent. Arafura Sea,64.5 km N of Croker Island, Northern Territory,10°17’S, 132°38’E, 65 m. Holotype AMSC174625; paratypes AMS C173679 (7). Caduluswoodhousae Lamprell & Healy, 1998. Indo-Pacific:northern Australia. 65-1380 m.woolacottae, Dentalium (Antalis) Colman, 1958: 142,fig. 4. Recent. Cronulla, New South Wales,Australia. Holotype AMS C21230; paratypes AMSC170642 (ex C21230) (1), AMS C32706 (2;Middle Harbour, Sydney, New South Wales,Australia). Dentalium woolacottae Colman, 1958.Indo-Pacific: eastern Australia. 9-40 m.† wymensis, Laevidentalium“?” Kulikov, 1967: 116,pl. 1, fig. 2. Permian. Vem River, Komi Republic,Russia. Holotype Geological Institute (St Petersbourg),8758/3.† xiphias, Dentalium Sharp & Pilsbry in Pilsbry &Sharp, 1898: 219. Eocene. Paris Basin (Parnes),France. Replacement name for Dentalium affineDeshayes, 1861 non Biondi, 1859. Objective juniorsynonym: Dentalium desmoulinsi Le Renard, 1994.Remark: a valid species of Dentalium (fide LeRenard 1995: 175; Le Renard & Pacaud 1995: 85,as “D. desmoulinsi”).yamakawai, Dentalium Yokoyama, 1927: 427, pl. 48,fig. 6. Pleistocene. Shinagawa, Tokyo [Japan].Lectotype UMT CM.23906 (designated by Taki &Oyama 1954: caption to pl. 45, as “holotype”).Junior synonym of Episiphon virgula (Hedley,1903) (fide Habe 1964: 28, as E. makiyamai).Remarks: reported from the Recent fauna of Japanby Nomura & Niino (1940: 63) and Nomura &Hatai (1940: 73, 74). Oyama (1973: 3) explained thelectotype designations of Taki & Oyama (1954).† yasilum, Dentalium Olsson, 1930: 72, pl. 12,figs 14-20. Eocene, Talara Formation. Yasila, Peru.Holotype PRI 24299; paratypes PRI 24297, PRI24298, PRI 24300-24303 (fide Brann & Kent1960: 323).† yatalensis, Cadulus (Dischides) Ludbrook, 1956: 4, 5,pl. 1, figs 3, 4. Pliocene. Weymouth’s Bore, DryCreek Sands, South Australia. Holotype SAM F15142; paratypes SAM (28). Remark: this specieswas misidentified as Cadulus mucronatus Tate, 1887by Ludbrook (1941).yateensis, Dischides Scarabino, 1995: 343, figs 146,147g. Recent. Yaté, New Caledonia, LAGON stn619, 22°03’S, 166°54’E, 27-42 m. Holotype andparatype MNHN. Dischides yateensis Scarabino,1995. Indo-Pacific: Philippines and New Caledonia.0-42 m.yokohamense, Dentalium Watson, 1879: 517 [Watson1886: 11, pl. 2, fig. 1]. Recent. Yokohama, Japan,Challenger stn 233, 34°39’N, 135°14’E, 8-14 fms[15-26 m]. Syntypes BMNH 1887.2.9.45-47 (3).Junior synonym of Dentalium octangulatumDonovan, 1803 (fide Habe 1964: 8).yokoyamai, Dentalium Makiyama, 1931: 44, pl. 1,fig. 1. Recent. Koshiba, Yokosuka City, Honshu,Pleistocene of Japan. Holotype KIU JC.310001(fide Hanzawa et al. 1961: 210). Fissidentaliumyokoyamai (Makiyama, 1931) (fide Kira 1959:106). Indo-Pacific: northern Australia to China Sea.56-504 m. Remark: this species was misidentified asDentalium complexum Dall, 1895 by Yokoyama(1920).† yotsukurensis, Dentalium Hirayama, 1955: 109-110,pl. 4, figs 26, 28. Upper Oligocene, Uchigo Group.Hirayama Locality A15, 1 km N of YoksukuraFishing Port, Fukushima Prefecture, Japan.Holotype GIUT 10233; paratypes GIUT 10208,10210 (Hirayama localities A28 and A45; fideHanzawa et al. 1961: 210).zanzibarense, Dentalium Plate, 1908a: 348, pl. 30,figs 35, 36. Recent. Zanzibar Channel, Valdiviastn 245, 05°28’S, 39°18’E, 463 m. Holotype ZMB61092 (fide Kilias 1995: 177). Compressidentaliumzanzibarense (Plate, 1908) (fide Scarabino 1995:262). Indo-Pacific: Africa to Madagascar. 463-1850 m.† zecaninus, Cadulus Laws, 1939: 503, pl. 63, fig. 18.Miocene. Pakaurangi Point, Kaipara Harbour, NewZealand. Holotype NZGS TM1374 (fide Keyes1972: 98). Gadila zecanina (Laws, 1939) (fideMaxwell 1992: 187).zeidleri, Laevidentalium Lamprell & Healy, 1998:114, figs 112E, 113F, 119, 120. Recent. 29 km SEof Scamander, Tasmania, Australia, 41°33’S,148°36’E, 122 m. Holotype SAM D 18903.Laevidentalium zeidleri Lamprell & Healy, 1998.Known from the type locality only.zelandicum, Dentalium Sowerby, 1860: 101, pl. 223,fig. 13. Recent. New Zealand. Type material notlocated in BMNH. Junior synonym: Dentaliumpacificum Hutton, 1873 (fide Suter 1913: 819).Fissidentalium zelandicum (Sowerby, 1860) (fideSuter 1913: 819). Indo-Pacific: New Zealand;Antarctica. 45-548 m. Remark: Dentalium opacum664ZOOSYSTEMA • 2004 • 26 (4)
Species-group names of Scaphopoda (Mollusca)Sowerby, 1829 may be a senior synonym for thisspecies (fide Dell 1957: 564).† zephyrinum, Dentalium Casey, 1903: 267.Oligocene. Red Bluff, Mississippi, USA. LectotypeUSNM Paleobiology 481672 (designated byMacNeil & Dockery 1984: 247, as “holotype”).† zingulai, Cadulus (Gadila) Hodgkinson, 1974: 18, textfig.8v, w; pl. 1, fig. 11, pl. 2, fig. 6a, b. Middle Eocene,Claibornian Stage, Cook Mountain Formation,Wheelock Member. Little Brazos Bluff, Brazos RiverValley, Brazos County, Texas, USA. Holotype USNMPaleobiology 180435; paratypes USNM Paleobiology180447, PRI 29227, ANSP 31505.zonatum, Dentalium “Orb.” Paetel, 1888: 594. “Jam.”[= Jamaica]. Nomen nudum. Remark: Pilsbry &Sharp (1898: 253) stated that this species “isunknown to us”.zonatus, Cadulus (Gadila) Boissevain, 1906: 74, pl. 6,fig. 57, text-fig. 37. Recent. Banda Sea, Indonesia,Siboga stn 214, 06°30’S, 121°55’E, 2796 m.Lectotype ZMA 3.06.102 (designated by Scarabino1995: 358) ; paralectotype ZMA 3.06.103(1; Arafura Sea, Siboga stn 254, 05°40’S, 132°26’E,310 m). Gadila zonata (Boissevain, 1906) (fideScarabino 1995: 358). Indo-Pacific: Indonesia toPhilippines. 195-2796 m.OTHER TAXA DESCRIBEDAS SCAPHOPODAThe list of non-scaphopod species erroneouslydescribed or assigned to this class, as compiled byPilsbry & Sharp (1898) is reproduced here, supplementedby names subsequently removed fromthe Recent Scaphopoda. We have included thetype species of the six genera – Entalium, Falcula,Hamulus, Pharetrium, Pyrgopolon and Spirodentaliumthat Pilsbry & Sharp (1898: 244-247)excluded from the Scaphopoda. However, wehave not included any species that were subsequentlydescribed in those six genera, since theydo not enter into homonymy with valid taxa ofthe Scaphopoda. The entry form is that of the listof genus-group and Recent species-group names.Detailed information on type localities, typematerial deposits and synonymy is only providedfor the names removed from Scaphopoda subsequentto Pilsbry & Sharp (1898).abbreviatum, Dentalium Deshayes, 1825: 352,pl. xviii, figs 21, 22 (“1864”: 199, pl. 3, figs 5-7).Eocene of Paris. “Serpula” heptagona Sowerby (fidePilsbry & Sharp 1898: 241) (Polychaeta).aciculatum, Dentalium Hall, 1860: 107. MiddleDevonian, Marcellus Shale and Portage Groups.Manilus, New York, USA. Coleolus aciculatus, transferredto Coleolus (phylum uncertain) by Hall(1879: 190), but it may as likely be a scaphopod(fide Pilsbry & Sharp 1898: 240).acus, Dentalium Eichwald, 1857: 584. Silurian[“Grauwacke”]. Putilowa [Poulkovo], near StPetersbourg, Russia; near Erras, Estonia.Criconarida (Cephalopoda: Mollusca) (fide Kiselev2001). Remarks: figured by Eichwald, 1860: 1062,pl. 40, fig. 10. Koken (1897: 214) remarked thatEichwald’s illustration showed an unusual irregularityin its curvature (“eine Unregelmässigkeit in derBiegung”), and questioned whether this species wasreferable to the Scaphopoda. Kiselev (2001) concludedthat this species was actually from theOrdovician, and belongs to the Cephalopoda.annulata, Dentalium McClelland, 1834: 629, pl. 35,figs 1-4. Ponar River, Kamaon, northern India.Remark: the biological nature of this taxon, whichappears to be a trace fossil, cannot be determined.annulatum, Dentalium Jay, 1850: 96 [ex Mighels MS]non Dentalium annulatum Gmelin, 1791. “Maine”.Nomen nudum. Remarks: Pilsbry & Sharp (1898:240) suggested that this taxon might be a Caecum(Gastropoda). We know of no basis for suggestingthat a nomen nudum could be referable to a knowngenus.antiquum, Dentalium Goldfuss, 1841: 2, pl. 166,fig. 2. Devonian? Eifel Mountain, Germany.Coleolus antiquum (Goldfuss, 1841) (fide Yochelson2002). Yochelson (2002) concluded that thisspecies was not a scaphopod, and suggested that itbe referred to Coleolus Hall, a calcareous “worm”tube.arietinum, Dentalium Müller, 1776: 236. Ditrupa arietina(Müller) (Polychaeta) (fide Pilsbry & Sharp1898: 242).beachportensis, Tesseracme Cotton & Ludbrook, 1938:220, pl. 12, fig. 2. Off Beachport, South Australia,201 m. Holotype SAM D 13339. Pennatulidrhachis (Cnidaria) (fide Lamprell & Healy 1998:172).belcheri, Prodentalium Nassichuk & Hodgkinson,1976: 1153-1156, pl. 1, figs 1-9. Early Permian,Assistance Formation. GSC Locality 246406, LyallRiver, Grinnell Peninsula, Devon Island, ArcticOcean, Canada. Holotype GSC 1377; numerousparatypes GSC 13776, 44794-44797 (Polychaeta?).Remark: Yancey (1978: 306) stated that this species“is not a species of Prodentalium, and probably nota scaphopod” and concluded that “it is probablethat it is not a molluscan species, but a calcareousworm tube”.billingsi, Salterella Safford, 1869: 289. Ordovician ofTennessee. Transferred to the Scaphopoda byZOOSYSTEMA • 2004 • 26 (4)665
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