Steiner G. & Kabat A. R.Dentalium annulatum Meyer, 1886 non Dentaliumannulatum Gmelin, 1791. Herein renamedDentalium parannulatum Steiner & Kabat, 2004.Dentalium asperum Brunet, 1995 non Dentalium asperumMichelotti, 1847. Herein renamed Dentaliumbruneti Steiner & Kabat, 2004.Dentalium bicarinatum Wanner, 1902 non Dentaliumbicarinatum Deshayes, 1825. Herein renamedDentalium wanneri Steiner & Kabat, 2004.Dentalium brevifissum Brugnone, 1876 non Dentaliumbrevifissum Deshayes, 1825. Herein renamedDentalium brugnonei Steiner & Kabat, 2004.Dentalium duodecimcostatum Brazier, 1877 nonDentalium duodecimcostatum d’Archiac, 1850.Herein renamed Dentalium healyi Steiner & Kabat,2004.Dentalium (Laevidentalium) ellipticum Koenen, 1892non Dentalium ellipticum J. Sowerby, 1814. Hereinrenamed Laevidentalium pseudellipticum Steiner &Kabat, 2004.Dentalium flavum Scarabino, 1995 non Dentaliumflavum Henderson, 1920. Herein renamed Dentaliumscarabinoi Steiner & Kabat, 2004.Dentalium hexagonum Van Ingen, 1905 nonDentalium hexagonum Gould, 1859. Hereinrenamed Dentalium vaningeni Steiner & Kabat,2004.Cadulus martini Scarabino, 1995 non Cadulus martiniFinlay, 1927. Herein renamed Cadulus scarabinoiSteiner & Kabat, 2004.Dentalium (Tesseracme) mexicanum Perrilliat, 1975non Dentalium mexicanum Girty, 1909. Hereinrenamed Dentalium perrilliatae Steiner & Kabat,2004.Dentalium minor Stephenson, 1952 non Dentaliumcaudani var. minor Locard, 1898. Herein renamedDentalium minorinum Steiner & Kabat, 2004.Dentalium polygonum Casey, 1903 non Dentaliumpolygonum Reuss, 1844. Herein renamed Dentaliumcaseyi Steiner & Kabat, 2004.APPENDIX 2Arguments for the taxonomic decisions.NEW GENERIC COMBINATIONSDentalium americanum Chenu, 1843 has previouslybeen placed in Dentalium (e.g., Pilsbry &Sharp 1897: 22; Henderson 1920: 27; Scarabino1975: 183). This <strong>species</strong> shows six primary ribscharacteristic for the genus Paradentalium,whereas Dentalium has eight to 12 primary ribs.Therefore, we have transferred this <strong>species</strong> toParadentalium.Dentalium peruvianum Dall, 1908 has the followingcharacters diagnostic <strong>of</strong> Fissidentalium:large (90 mm), almost straight shell with slightlydepressed anterior aperture; a high number <strong>of</strong>primary ribs (about 26); a long <strong>and</strong> wide slit onthe ventral side <strong>of</strong> the apex. In agreement with V.Scarabino (in litt. 6.III.2000), we have transferredthis <strong>species</strong> to Fissidentalium.Dentalium cancellatum Sowerby, 1860 shows sixto eight primary ribs (intercalating towards theanterior aperture) intersected by raised transversestriae giving the shell a cancellated appearancetypical for Coccodentalium. Similar patterns <strong>of</strong>sculpture are also present in some <strong>species</strong> <strong>of</strong>Fissidentalium <strong>and</strong> Compressidentalium, both <strong>of</strong>which have more than 12 primary ribs. We,therefore, transferred this <strong>species</strong> to Coccodentalium.Cadulus (Platyschides) foweyensis Henderson,1920 has four apical lobes, two lateral <strong>and</strong> oneeach dorsal <strong>and</strong> ventral, separated by distinctnotches. This is typical for the genus Polyschides.We, therefore, transferred this <strong>species</strong> toPolyschides.Cadulus (Platyschides) nitidus Henderson, 1920has four apical lobes, two lateral <strong>and</strong> one eachdorsal <strong>and</strong> ventral, separated by distinct notches.This is typical for the genus Polyschides. We,therefore, have transferred this <strong>species</strong> toPolyschides.Cadulus poculum Dall, 1889 has been transferredto the genus Polyschides because <strong>of</strong> the position <strong>of</strong>the maximum diameter being in the anteriorthird <strong>of</strong> the shell <strong>and</strong> the lateral projections at theapex indicating four apical lobes. Henderson710ZOOSYSTEMA • 2004 • 26 (4)
Species-<strong>group</strong> <strong>names</strong> <strong>of</strong> <strong>Scaphopoda</strong> (Mollusca)(1920: 108, 109) listed this <strong>species</strong> in his subgenusPlatyschides that was synonymized withGadila by Scarabino (1995). Although the placementinto Gadila is possible due to the inadequatelypreserved apical features, the stronglyoblique anterior aperture resembles that <strong>of</strong> severalPolyschides <strong>species</strong>.Cadulus (Platyschides) portoricensis Henderson,1920 has four apical lobes typical for the genusPolyschides. We, therefore, have transferred this<strong>species</strong> to Polyschides.Siphodentalium teres Jeffreys, 1883 has beenremoved from Siphonodentalium because it hasneither more than four apical lobes nor an anteriorconstriction, which is diagnostic <strong>of</strong>Siphonodentalium. Because <strong>of</strong> the long <strong>and</strong> slightlytapering shell <strong>and</strong> the radula morphology(polygonal, unicuspidate rhachis; lateral toothwith three cusps, the posterior two being closer toeach other <strong>and</strong> separated from the anterior cuspby a nodulous ridge), we have transferred this<strong>species</strong> to Pulsellum. The two lateral lobes atthe shell apex, however, are unique among the<strong>Scaphopoda</strong>.NEW SYNONYMIESThe following taxa have previously been recognizedas a valid <strong>species</strong>, or as infrasubspecific taxa,<strong>of</strong> Fissidentalium: Dentalium ergasticum Fischer,1883 (Pilsbry & Sharp 1897: 74, 75) <strong>and</strong> itsforms D. e. var. albida Locard, 1898, D. e. var.elongata Locard, 1898, D. e. var. major Locard,1896, D. e. var. minor Locard, 1898; D. milneedwardsiLocard, 1897; D. scamnatum Locard,1897 (Pilsbry & Sharp 1897: 79) <strong>and</strong> its formsD. s. var. alba Locard, 1898, D. s. var. minorLocard, 1898, D. s. var. subrecta Locard, 1898.However, these taxa fall within the range <strong>of</strong> variation<strong>of</strong> Fissidentalium capillosum (Jeffreys, 1877),as Bernard Métivier (MNHN; unpublished, pers.comm.) has demonstrated by his comparison <strong>of</strong>available type material <strong>and</strong> additional material.The synoymy <strong>of</strong> D. ergasticum Fischer, 1883,D. milneedwardsi Locard, 1897, <strong>and</strong> D. scamnatumLocard, 1897 with Fissidentalium capillosum(Jeffreys, 1877) is, therefore, credited toB. Métivier. Note that this synonymy changesthe name <strong>of</strong> the type <strong>species</strong> <strong>of</strong> the genusFissidentalium from Fissidentalium ergasticum toFissidentalium capillosum.Dentalium caudani Locard, 1896 <strong>and</strong> its formsD. c. var. parfaiti Locard, 1898, D. c. var. intermediaLocard, 1898, <strong>and</strong> D. c. var. minor Locard,1898 have previously been recognized as a valid<strong>species</strong>, or as infrasubspecific taxa, <strong>of</strong> Laevidentalium(Pilsbry & Sharp 1897: 104).However, these taxa fall within the range <strong>of</strong> variation<strong>of</strong> Fissidentalium c<strong>and</strong>idum (Jeffreys, 1877),as Bernard Métivier (MNHN; unpublished, pers.comm.) has demonstrated by his comparison <strong>of</strong>available type material <strong>and</strong> additional material.The synonymy <strong>of</strong> D. caudani Fischer, 1883 withF. c<strong>and</strong>idum (Jeffreys, 1877) is, therefore, creditedto B. Métivier.Dentalium exuberans Locard, 1897 <strong>and</strong> its formD. e. var. minor Locard, 1898 have previouslybeen recognized as a valid <strong>species</strong>, or as infrasubspecifictaxon, <strong>of</strong> Fissidentalium (Pilsbry & Sharp1897: 78, 79). However, shell shape <strong>and</strong> sculpture<strong>of</strong> these taxa are identical with the previouslydescribed F. paucicostatum (Watson, 1879), asBernard Métivier (MNHN; unpublished, pers.comm.) has demonstrated by his comparison <strong>of</strong>available type material <strong>and</strong> additional material.The synonymy <strong>of</strong> D. exuberans Locard, 1897with F. paucicostatum (Watson, 1879) is, therefore,credited to B. Métivier.Dentalium (Fustiaria) inversum Cossmann, 1915is placed in tentative synonymy with Fustiariamontense (Briart & Cornet, 1889) because it differsonly in the dorsal position <strong>of</strong> the apical slit.Cossmann (1915: 7) remarked that the type specimenmay be only an inverted specimen <strong>of</strong>Fustiaria montense. Dorsal displacement <strong>of</strong> the slitinfrequently occurs in other <strong>species</strong> <strong>of</strong> the genus(G. S. pers. obs.) <strong>and</strong> is, thus, not considered sufficientto separate a <strong>species</strong>, especially since thedescription was based on a single specimen only.Dentalium tessellatum Scarabino, 1995 matchesthe description <strong>and</strong> illustration <strong>of</strong> Dentaliumhabutae Kuroda & Kikuchi, 1933 in all details <strong>of</strong>shell shape <strong>and</strong> sculpture (V. Scarabino pers.comm., <strong>and</strong> in litt. 14.I.1997). Dentaliumhabutae is considered a valid <strong>species</strong> <strong>of</strong>ZOOSYSTEMA • 2004 • 26 (4)711
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