Catalog of species-group names of Recent and fossil Scaphopoda ...

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Steiner G. & Kabat A. R.Dentalium annulatum Meyer, 1886 non Dentaliumannulatum Gmelin, 1791. Herein renamedDentalium parannulatum Steiner & Kabat, 2004.Dentalium asperum Brunet, 1995 non Dentalium asperumMichelotti, 1847. Herein renamed Dentaliumbruneti Steiner & Kabat, 2004.Dentalium bicarinatum Wanner, 1902 non Dentaliumbicarinatum Deshayes, 1825. Herein renamedDentalium wanneri Steiner & Kabat, 2004.Dentalium brevifissum Brugnone, 1876 non Dentaliumbrevifissum Deshayes, 1825. Herein renamedDentalium brugnonei Steiner & Kabat, 2004.Dentalium duodecimcostatum Brazier, 1877 nonDentalium duodecimcostatum d’Archiac, 1850.Herein renamed Dentalium healyi Steiner & Kabat,2004.Dentalium (Laevidentalium) ellipticum Koenen, 1892non Dentalium ellipticum J. Sowerby, 1814. Hereinrenamed Laevidentalium pseudellipticum Steiner &Kabat, 2004.Dentalium flavum Scarabino, 1995 non Dentaliumflavum Henderson, 1920. Herein renamed Dentaliumscarabinoi Steiner & Kabat, 2004.Dentalium hexagonum Van Ingen, 1905 nonDentalium hexagonum Gould, 1859. Hereinrenamed Dentalium vaningeni Steiner & Kabat,2004.Cadulus martini Scarabino, 1995 non Cadulus martiniFinlay, 1927. Herein renamed Cadulus scarabinoiSteiner & Kabat, 2004.Dentalium (Tesseracme) mexicanum Perrilliat, 1975non Dentalium mexicanum Girty, 1909. Hereinrenamed Dentalium perrilliatae Steiner & Kabat,2004.Dentalium minor Stephenson, 1952 non Dentaliumcaudani var. minor Locard, 1898. Herein renamedDentalium minorinum Steiner & Kabat, 2004.Dentalium polygonum Casey, 1903 non Dentaliumpolygonum Reuss, 1844. Herein renamed Dentaliumcaseyi Steiner & Kabat, 2004.APPENDIX 2Arguments for the taxonomic decisions.NEW GENERIC COMBINATIONSDentalium americanum Chenu, 1843 has previouslybeen placed in Dentalium (e.g., Pilsbry &Sharp 1897: 22; Henderson 1920: 27; Scarabino1975: 183). This species shows six primary ribscharacteristic for the genus Paradentalium,whereas Dentalium has eight to 12 primary ribs.Therefore, we have transferred this species toParadentalium.Dentalium peruvianum Dall, 1908 has the followingcharacters diagnostic of Fissidentalium:large (90 mm), almost straight shell with slightlydepressed anterior aperture; a high number ofprimary ribs (about 26); a long and wide slit onthe ventral side of the apex. In agreement with V.Scarabino (in litt. 6.III.2000), we have transferredthis species to Fissidentalium.Dentalium cancellatum Sowerby, 1860 shows sixto eight primary ribs (intercalating towards theanterior aperture) intersected by raised transversestriae giving the shell a cancellated appearancetypical for Coccodentalium. Similar patterns ofsculpture are also present in some species ofFissidentalium and Compressidentalium, both ofwhich have more than 12 primary ribs. We,therefore, transferred this species to Coccodentalium.Cadulus (Platyschides) foweyensis Henderson,1920 has four apical lobes, two lateral and oneeach dorsal and ventral, separated by distinctnotches. This is typical for the genus Polyschides.We, therefore, transferred this species toPolyschides.Cadulus (Platyschides) nitidus Henderson, 1920has four apical lobes, two lateral and one eachdorsal and ventral, separated by distinct notches.This is typical for the genus Polyschides. We,therefore, have transferred this species toPolyschides.Cadulus poculum Dall, 1889 has been transferredto the genus Polyschides because of the position ofthe maximum diameter being in the anteriorthird of the shell and the lateral projections at theapex indicating four apical lobes. Henderson710ZOOSYSTEMA • 2004 • 26 (4)
Species-group names of Scaphopoda (Mollusca)(1920: 108, 109) listed this species in his subgenusPlatyschides that was synonymized withGadila by Scarabino (1995). Although the placementinto Gadila is possible due to the inadequatelypreserved apical features, the stronglyoblique anterior aperture resembles that of severalPolyschides species.Cadulus (Platyschides) portoricensis Henderson,1920 has four apical lobes typical for the genusPolyschides. We, therefore, have transferred thisspecies to Polyschides.Siphodentalium teres Jeffreys, 1883 has beenremoved from Siphonodentalium because it hasneither more than four apical lobes nor an anteriorconstriction, which is diagnostic ofSiphonodentalium. Because of the long and slightlytapering shell and the radula morphology(polygonal, unicuspidate rhachis; lateral toothwith three cusps, the posterior two being closer toeach other and separated from the anterior cuspby a nodulous ridge), we have transferred thisspecies to Pulsellum. The two lateral lobes atthe shell apex, however, are unique among theScaphopoda.NEW SYNONYMIESThe following taxa have previously been recognizedas a valid species, or as infrasubspecific taxa,of Fissidentalium: Dentalium ergasticum Fischer,1883 (Pilsbry & Sharp 1897: 74, 75) and itsforms D. e. var. albida Locard, 1898, D. e. var.elongata Locard, 1898, D. e. var. major Locard,1896, D. e. var. minor Locard, 1898; D. milneedwardsiLocard, 1897; D. scamnatum Locard,1897 (Pilsbry & Sharp 1897: 79) and its formsD. s. var. alba Locard, 1898, D. s. var. minorLocard, 1898, D. s. var. subrecta Locard, 1898.However, these taxa fall within the range of variationof Fissidentalium capillosum (Jeffreys, 1877),as Bernard Métivier (MNHN; unpublished, pers.comm.) has demonstrated by his comparison ofavailable type material and additional material.The synoymy of D. ergasticum Fischer, 1883,D. milneedwardsi Locard, 1897, and D. scamnatumLocard, 1897 with Fissidentalium capillosum(Jeffreys, 1877) is, therefore, credited toB. Métivier. Note that this synonymy changesthe name of the type species of the genusFissidentalium from Fissidentalium ergasticum toFissidentalium capillosum.Dentalium caudani Locard, 1896 and its formsD. c. var. parfaiti Locard, 1898, D. c. var. intermediaLocard, 1898, and D. c. var. minor Locard,1898 have previously been recognized as a validspecies, or as infrasubspecific taxa, of Laevidentalium(Pilsbry & Sharp 1897: 104).However, these taxa fall within the range of variationof Fissidentalium candidum (Jeffreys, 1877),as Bernard Métivier (MNHN; unpublished, pers.comm.) has demonstrated by his comparison ofavailable type material and additional material.The synonymy of D. caudani Fischer, 1883 withF. candidum (Jeffreys, 1877) is, therefore, creditedto B. Métivier.Dentalium exuberans Locard, 1897 and its formD. e. var. minor Locard, 1898 have previouslybeen recognized as a valid species, or as infrasubspecifictaxon, of Fissidentalium (Pilsbry & Sharp1897: 78, 79). However, shell shape and sculptureof these taxa are identical with the previouslydescribed F. paucicostatum (Watson, 1879), asBernard Métivier (MNHN; unpublished, pers.comm.) has demonstrated by his comparison ofavailable type material and additional material.The synonymy of D. exuberans Locard, 1897with F. paucicostatum (Watson, 1879) is, therefore,credited to B. Métivier.Dentalium (Fustiaria) inversum Cossmann, 1915is placed in tentative synonymy with Fustiariamontense (Briart & Cornet, 1889) because it differsonly in the dorsal position of the apical slit.Cossmann (1915: 7) remarked that the type specimenmay be only an inverted specimen ofFustiaria montense. Dorsal displacement of the slitinfrequently occurs in other species of the genus(G. S. pers. obs.) and is, thus, not considered sufficientto separate a species, especially since thedescription was based on a single specimen only.Dentalium tessellatum Scarabino, 1995 matchesthe description and illustration of Dentaliumhabutae Kuroda & Kikuchi, 1933 in all details ofshell shape and sculpture (V. Scarabino pers.comm., and in litt. 14.I.1997). Dentaliumhabutae is considered a valid species ofZOOSYSTEMA • 2004 • 26 (4)711
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Catalog of species-group names of Recent and fossil Scaphopoda ...

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