Catalog of species-group names of Recent and fossil Scaphopoda ...

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Steiner G. & Kabat A. R.rejected Noetling’s claim that his Burma specimenswere conspecific with Dentalium junghuhnidescribed from Java, Indonesia. Noetling (1901:250) claimed that his Burma specimens of“junghuhni” were “absolutely identical with”Dentalium magnificum E. A. Smith, 1896, a Recentspecies from the Indian Ocean. Martin’s subsequentconclusion that the Burma specimens were notreferable to Dentalium junghuhni means thatDentalium magnificum is no longer a junior synonym,and indeed may now be a senior synonym ofDentalium noetlingi.† normanianum, Dentalium d’Orbigny, 1850 (volume2): 46. Jurassic. Villerville, France. Remark:this species was misidentified as Dentalium nitens byEudes-Deslongchamp (1842).† notabile, Dentalium Eichwald, 1857: 583. Silurian[“Grauwackenkalksteine”]. Pulkowa, Russia.Remark: figured by Eichwald, 1860: 1061, pl. 40,fig. 9; 1868: 800, 801, pl. 28, fig. 1.novaehollandiae, Dentalium Chenu, 1843: pl. 6,fig. 14; 1852: 5. Recent. Australia. Type materialnot located. Graptacme? novaehollandiae (Chenu,1843) (fide Pilsbry & Sharp 1897: 93). No informationon geographic or bathymetric distributionavailable.† novaki, Dentalium Koenen, 1892: 978-980, pl. 59,figs 7, 8. Oligocene. Morigny, France. Remark:Merklin & Goncharova (1967: 118) listed novaki asa junior synonym of acutum Hébert, but Janssen(1978b: 199) treated it as a valid species of Fissidentalium.† novaresei, Dentalium (Plagioglypta) de Stefani, 1917:42, 43, pl. 4, fig. 6. Carboniferous. Elba, Italy.† novemcinctum, Dentalium “?” Sacco, 1897: 96, 97,pl. 7, fig. 97 [as a nomen nudum in Sacco 1896: 97].Oligocene. Sassello, Italy. Holotype MIGTBS.106.01.045 (fide Ferrero Mortara et al. 1984:303, pl. 56, fig. 1). Remark: Bonci et al. (2000:206) concluded that this was a valid species, referableto Dentalium (Dentalium).novemcostatum, Dentalium Lamarck, 1818: 344.Recent. Around La Rochelle, France [Atlantic].Possible syntypes MHNG 1081/54 (5; fide Y.Finet, in litt. 16.X.1996). Antalis novemcostata(Lamarck, 1818) (fide Pilsbry & Sharp 1897: 51).Eastern Atlantic: Europe. 20-300 m.noviluna, see novilunatus.novilunatus, Cadulus (Platyschides) Kira, 1959: 104,pl. 40, fig. 2 [nomen nudum in Habe 1953: 298,and in Kira 1954: 80, pl. 40, fig. 2; see Bieler &Petit 1990: 141]. Recent. Shikoku, Japan. Typematerial not located. Junior synonym of Gadila virginalis(Boissevain, 1906) (fide Habe 1963: 279).Remarks: although Habe (1963: 279, 1964: 49,1977: 342) and Habe & Kosuge (1964: 12) includedG. novilunata in the synonymy of virginalis,Scarabino (1995: 358) stated that he “... prefer[s] tokeep them separate pending further study”. Gadilanoviluna [Habe 1977: 342] is an incorrect subsequentspelling.† novum, Dentalium Chenu, 1843: pl. 6, fig. 23;1850: 5. Type locality not specified (Tertiary, fidePilsbry & Sharp 1898: 210).† nudum, Dentalium Zekeli, 1852: 118, pl. 24, figs 1,2. Jurassic. Edelbachgraben, Gosautal, Austria.numatai, Dentalium (Fustiaria) Hirase, 1931: 139,pl. 3, fig. 11. Recent. Osumi and Oshima [Japan]. Typematerial not located. Junior synonym of Fustiarianipponica (Yokoyama, 1922) (fide Habe 1963: 267).† numerosa, Plagioglypta Yoo, 1988: 248, figs 148-151.Early Carboniferous, Late Tournaisian, DangarfieldFormation. Locality A-3, 2.6 km SW of Gundy,Upper Hunter, New South Wales, Australia. HolotypeAMS F.61996; paratypes AMS F.61997 (20).numerosum, Dentalium Dall, 1895: 686. Recent. Fromthe Galapagos to California. Syntypes LACM 1755(near Isla Cedros, Pacific coast Baja California,México, 28°12’N, 115°09’W, 80 m), SBMNH35016, USNM 87559, USNM 203102, USNM600413 (all from Baja California Norte, near IslaCedros, Albatross stn 2823, 24°18’N, 110°22’W,26 fms [48 m]). Junior synonym of Dentalium oerstediiMörch, 1861 (fide Pilsbry & Sharp 1897: 25).† nunomae, Dentalium Takeda, 1953: 62, 63, pl. 4,fig. 8, pl. 5, fig. 12. Oligocene, Poronai Formation.Hokkaido University Locality T268, southernSakhalin Island, Russia. Holotype HokkaidoUniversity No. 11090; paratype HokkaidoUniversity No. 11089 (Locality T211).† nutans, Cadulus Böhm, 1891: 70, pl. 4, fig. 26.Cretaceous. Höpfling, near Siegsdorf, southernBavaria, Germany.nutans, Dentalium Kner, 1850: 23, 24, pl. 4, figs 10,10a. Cretaceous. Nagórzany, Ukraine. Junior synonymof Dentalium decussatum J. Sowerby, 1814(fide Pilsbry & Sharp 1898: 221).nysti, Dentalium d’Orbigny, 1852 (volume 3): 18.Tertiary, Falunian. Klein-Spauwen, Hoesselt, Lethen,Belgium. Junior synonym of Dentalium acutumHébert, 1849 (fide Pilsbry & Sharp 1898: 197).Remark: this species was misidentified as Dentaliumgrande Deshayes, 1825 by Nyst (1843).nysti, Dentalium Van den Binkhorst, 1861: 61, pl. 6,fig. 2a-2c. Cretaceous. Limbourg, Belgium. Juniorhomonym of Dentalium nysti d’Orbigny, 1852.Replacement name: Dentalium binkhorsti Pilsbry &Sharp, 1898.obesus, Cadulus Watson, 1879: 527, 528 [Watson1886: 22, pl. 3, fig. 8]. Recent. Culebra Island[Lesser Antilles], Challenger stn 24, 18°38’30”N,65°05’30”W, 390 fms [712 m]. Syntypes BMNH1887.2.9.84-87 (labelled “in advanced stage ofdecay; Palmer 1971”). Cadulus obesus Watson,1879. Western Atlantic, Caribbean. 365-712 m.† obliquatus, Cadulus Koenen, 1892: 990, pl. 59, figs19, 20. Lower Oligocene. Lattorf, Calbe, Atzendorf,and Unseburg, Germany.622ZOOSYSTEMA • 2004 • 26 (4)
Species-group names of Scaphopoda (Mollusca)† oblongum, Dentalium Richardson, 1906: 585,pl. 45, fig. 3. Jurassic, Pliensbachian. Railwaycutting,Dixton West, near Gotherington, nearCheltenham, England, United Kingdom. Remark:Engeser & Riedel (1992: 38) stated that “Thisspecies cannot be attributed to any known scaphopodgenus and therefore a new genus should beproposed”.obnutus, see obrutus.† obrutus, Gadus Conrad, 1870a: 227. Emendationfor Gadus “obnutus” Conrad, 1869b. Cretaceous.Haddonfield, New Jersey, USA. Syntype ANSP19502. Cadulus obrutus (Conrad, 1870) (fidePilsbry & Sharp 1898: 238).obscurum, Dentalium gouldii var. Dall, 1889: 425,pl. 27, fig. 4. Recent. Off Barbados, Lesser Antilles,Blake stn 299, 13°05’N, 59°39’40”W, 140 fms[256 m]. Lectotype USNM 95351 (designated byHenderson 1920: 32); paralectotypes USNM887460, MCZ 7695, MCZ 7708 (erroneouslystated to be the holotype by Turner 1955: 319).Dentalium obscurum Dall, 1889 (fide Henderson1920: 32). Western Atlantic: Caribbean and Brazil.146-1540 m.† obsoletum, Dentalium Hall, 1858: 724, pl. 29,figs 16, 17. Carboniferous. Iowa, USA. SyntypeFM UC.10722 (fide Forney & Nitecki 1976: 11).Objective junior synonyms: Dentalium sublaeveHall, 1877; Prodentalium fredericae Engeser &Riedel, 1992. Junior synonym: Dentalium acutisulcatumGurley, 1883 (fide Yancey 1978: 307).Prodentalium obsoletum (Hall, 1858) (fide Engeser& Riedel 1992: 41). Remarks: this is a valid nameand not a junior secondary homonym of Dentalitesobsoletum Schlotheim, 1832, which is a nomennudum. The replacement names, Dentalium sublaeveHall in Miller, 1877 (being itself a juniorhomonym of Dentalium sublaeve Cocconi, 1874)and Prodentalium fredericae Engeser & Riedel, 1992are, therefore, unnecessary.obsoletum, Dentalium Doderlein, 1864: 97 [15].Nomen nudum. Miocene. Monte Gibio (Modena),Santo Agata (Tortona) and Vigoleno (Piacenza),Italy. Junior homonym of Dentalium obsoletumHall, 1858. See Dentalium dentalis var. sublaevisCocconi, 1874.obsoletus, Dentalites Schlotheim, 1813: 100; 1832: 67.Tonna, Sachsenberg, Germany. Nomen nudum.obtusum, Dentalium Qi & Ma, 1989: 114, fig. 3.Recent. Zhejiang Province, China. Type in IOAS.Dentalium obtusum Qi & Ma, 1989. Indo-Pacific:China Sea. 0-5 m.occidentale, Dentalium Stimpson, 1851b: 28. Recent.Deep water on the coast of Maine, and in MassachusettsBay [Western Atlantic]. Type material presumablylost (fide Johnson 1976: 66). Juniorsynonyms: Dentalium abyssorum M. Sars, 1859 (fidePilsbry & Sharp 1897: 47); Dentalium cinerascensAnton, 1838 (fide Pilsbry & Sharp 1897: 48, as tentativesenior(!) synonym); Dentalium aenigmaticumJordan, 1895 (fide Friele & Grieg 1901: 49).Antalis occidentalis (Stimpson, 1851) (fide Pilsbry& Sharp 1897: 47). North Atlantic. 36-1823 m.Remarks: Emerson in Turgeon (1998: 54) listedthis species in the synonymy of Antalis entalis(Linnaeus, 1758); further study of the taxonomy ofthese species is needed. This species was misidentifiedas Dentalium dentale by Gould (1841).occidentale, Siphonodentalium (Pulsellum) Henderson,1920: 93, pl. 16, fig. 4. Recent. Off Martha’sVineyard [Massachusetts], USBF stn 871,40°05’39”N, 70°23’52”W, 86 fms [157 m]. HolotypeUSNM 38702; paratype MCZ 186823 (2).Pulsellum occidentale (Henderson, 1920). WesternAtlantic: Massachusetts, USA. 157-1794 m.occiduus, Cadulus Verco, 1911b: 218, pl. 26, fig. 7.Recent. Geographe Bay, off Bunbury, WestAustralia, 15 fms [27 m]. Holotype SAM D 13759.Cadulus occiduus Verco, 1911. Indo-Pacific: westernAustralia. 27-2900 m. Remark: the 7 paratypescould not be located.octangulatum, Dentalium Donovan, 1803: pl. 162.Recent. Japan. Neotype BMNH 1952.2.23.1 (designatedby Ludbrook 1954: 97). Junior synonyms:Dentalium octogonum Lamarck, 1818; Dentaliumstriatulum Turton, 1819 (fide Pilsbry & Sharp1897: 17); Dentalium yokohamense Watson, 1879;Dentalium japonicum Dunker, 1877 (fide Habe1964: 8). Dentalium octangulatum Donovan,1803. Indo-Pacific: Red Sea to Japan and NewGuinea. 5-1380 m. Remarks: the original typelocality was “near Lelant, Cornwall, England”.However, the type material was mislocalized, andLudbrook’s designation of a neotype from Japanfixes the type locality as Japan (ICZN 1999: article76.3).† octevillense, Dentalium Cossmann, 1899b: 329,pl. 2, fig. 8. Cretaceous, Cenomanian. Octeville,Seine-Inférieure [Seine-Maritime], France.octocostatum, Dentalium Fraas, 1867. Not a scaphopod.See Other taxa described as Scaphopoda.Remark: Blanckenhorn (1927: 123, 124, pl. 1,figs 2-8) considered this to be a valid species ofDentalium, but he did not discuss the contraryconclusion of Pilsbry & Sharp (1898: 245).octocostatum, Dentalium Ihering, 1897: 266, 267,pl. 4, fig. 16. Tertiary, Santa Cruz Formation.Yegua Quemada, Patagonia, Argentina. Juniorhomonym of Dentalium octocostatum Fraas, 1867[Polychaeta]. Replacement name: Dentalium octocostellatumPilsbry & Sharp, 1898.† octocostellatum, Dentalium Pilsbry & Sharp, 1898:211. Tertiary, Santa Cruz Formation. YeguaQuemada, Patagonia, Argentina. Replacementname for Dentalium octocostatum Ihering, 1897 nonFraas, 1867 (Polychaeta). Junior synonym:Dentalium gallardoi Rovereto, 1921 (fide Feruglio1949b: 203, footnote).ZOOSYSTEMA • 2004 • 26 (4)623
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