Catalog of species-group names of Recent and fossil Scaphopoda ...

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Steiner G. & Kabat A. R.Palestine. Polychaeta (fide Pilsbry & Sharp 1898:245). Remark: Blanckenhorn (1927: 123, 124, pl.1, figs 2-8) considered this to be a valid species ofDentalium, but he did not discuss the contraryconclusion of Pilsbry & Sharp.onyx, Hamulus Morton, 1834: 73, 74, pl. 2, fig. 8,pl. 16, fig. 5. Cretaceous. Lynch’s Creek, SouthCarolina and Erie, Alabama, USA. Polychaeta (fidePilsbry & Sharp 1898: 245).opiparium, Quasidentalium Shimansky, 1974: 135,figs 2a-2c. Lower Carboniferous, Donets Basin[Russia]. Coleolus (fide Yochelson 1999: 63-65).osceola, Spirodentalium Walcott, 1890: 271, 272,pl. 20, fig. 12. Upper Cambrian, Potsdam Terrane.Osceola Mills, Wisconsin, USA. Lectotype USNMPaleobiology 23853a (selected by Yochelson 1987:66, 67, fig. 1.4); two paralectotypes USNMPaleobiology 23853b, 23853c. Remark: Yochelson(1987) concluded that this taxon was actually anopen-coiled gastropod, which he referred to theEuomphaloidea.pellucidum, Dentalium Gmelin, 1791: 3738 [citingSchröter 1783-1786: 529]. Oceano septentrionali[northern ocean]. Polychaeta (fide Pilsbry & Sharp1898: 243, pl. 6, fig. 17).planum, Dentalium J. Sowerby, 1814: 179, pl. 79,fig. 1. Bognor, England. Polychaeta (fide Pilsbry &Sharp 1898: 243).pusillum, Dentalium Philippi, 1836: 245. Panormi,Catania [Sicily]. “Has been referred to Dischidespolitus, but it may be a Ditrupa” (Pilsbry & Sharp1898: 243).pygmaeus, Dentalium Defrance, 1819a: 71. Caecum(Gastropoda) (fide Pilsbry & Sharp 1898: 241).radicula, Dentalium Lamarck, 1818: 345. Eocene.Grignon, France. Polychaeta (fide Pilsbry & Sharp1898: 243).rudis, ?Dentalium Gabb, 1873: 244. San Domingo[Dominican Republic]. Polychaeta (fide Pilsbry &Sharp 1898a: 474, pl. 10, figs 4, 8; Pilsbry & Sharp1898: 243).rugosum, Dentalium Müller, 1851: 6, pl. 3, fig. 2.Cretaceous. Lusberg, Germany. Gastrochaena(Bivalvia) (fide Holzapfel 1888: 179). Remarks:Pilsbry & Sharp (1898: 227) initially suggested thatthis species might belong to the Serpulidae[Polychaeta] or Teredinidae [Bivalvia]; in theiraddendum, Pilsbry & Sharp (1898: 255) thennoted that Holzapfel (1888) had already placed thistaxon in the Bivalvia.rugosum, Entalium Defrance, 1819b: 518. Cretaceous.Saint-Pierre, near Maastricht, Netherlands.Polychaeta (fide Pilsbry & Sharp 1898: 245).septangulare, Dentalium Fleming, 1825: 240, pl. 9,fig. 1. Belfast [Ireland]. Probably “Serpula” heptagonaSowerby (Polychaeta) (fide Pilsbry & Sharp1898: 243).septemcostatum, Dentalium Abich in Trautschold,1859: 314, 315, pl. 6, fig. 5. Eocene and Oligoceneof Armenia. Probably “Serpula” heptagona Sowerby(Polychaeta) (fide Pilsbry & Sharp 1898: 244).serratum, Dentalium Pictet & Roux, 1849: 150, pl. 27,fig. 12a, b. Cretaceous, Bouches-du-Rhône, France.Probably belongs to Hamulus (Polychaeta) (fidePilsbry & Sharp 1898: 244).sexcarinatum, Dentalium Goldfuss, 1841: 4, pl. 166,fig. 12. Cretaceous. Friedland, Silesia (Pomerania?),Germany; St Petri, Maastricht, Netherlands.Polychaeta (fide Pilsbry & Sharp 1898: 244).sexradiatum, Dentalium Goldfuss – Pilsbry & Sharp1898: 244. “Craie super. Maestricht”. Nomennudum. May be an incorrect subsequent spelling ofsexcarinatum Goldfuss, 1844 (fide Pilsbry & Sharp1898: 244). Remark: Pilsbry & Sharp (1898: 244)state that they “have seen no work in which thisspecies is described”.sowerbyi, Dentalium Michelotti, 1847: 145. Miocene,Italy. Junior homonym of Dentalium sowerbyiGuilding, 1834. Polychaeta (fide Pilsbry & Sharp1898: 244).spinulosum, Dentalium Miller, MSS – Pilsbry & Sharp(1898: 241) cited this name in the synonymy of.Hamites spinulosus J. Sowerby, 1818 (Cephalopoda).spirale, Dentalium Risso, 1826b: 125; 1826b: 401.(Fossil). France. Type material presumed lost (fideArnaud 1978: 135). Polychaeta. Remark: Arnaud(1978: 135) noted that Risso’s unpublished illustrationof the type specimen showed spiral ornamentationthat is unknown in the Scaphopoda.strangulatum, Dentalium Deshayes, 1825: 372, 373,pl. 16, fig. 28. Australian Sea?, fossil in England,France, and Italy. Polychaeta (fide Pilsbry & Sharp1898: 244). Dentalium strangulosum “Deshayes”[Gümbel, 1861: 604] is an incorrect subsequentspelling.strangulosum, see strangulatum.“subcarinatum”, Dentalium “Münster in Goldfuss” –Ryckholt 1851. May be an incorrect subsequentspelling of Dentalium sexcarinatum Münster inGoldfuss, 1844 (fide Pilsbry & Sharp 1898: 244).subtilis, Cadulus (Gadila) Plate, 1908a: 360, pl. 30,fig. 48. Recent. Near Dar-es-Salaam, Tanzania,Valdivia stn 244, 5°55.8’S, 39°1.2’E, 50 m. LectotypeZMB 61108a (designated by Kilias 1995:176); paralectotypes ZMB 61108b (1), ZMB63842 (4). Remarks: Jaeckel (1932: 310) remarkedupon certain similarities with Ditrupa (Polychaeta),and Kilias (1995: 176) parenthetically remarkedthat this taxon is probably Ditrupa. Plate’s illustrationindeed resembles a worm tube, and the taxonmay have to be removed from the Scaphopoda afterexamination of the types.subulatum, Dentalium Deshayes, 1825: 373, pl. 16,fig. 29. Type locality not specified. Polychaeta (fidePilsbry & Sharp 1898: 244).tiwhana, Dentalium Dell, 1953: 48, figs 17, 22.Chatham Rise, New Zealand, 361 m. Serpula crenata668ZOOSYSTEMA • 2004 • 26 (4)
Species-group names of Scaphopoda (Mollusca)(Ehlers, 1908) (Polychaeta, Serpulida) (fide tenHove & Smith 1990: 107; fide Dell 1995: 21).trachea, Dentalium Montagu, 1803: 497, pl. 14,fig. 10. Caecum (Gastropoda) (fide Pilsbry & Sharp1898: 241).tricostatum, Dentalium Goldfuss, 1841: 3, 4, pl. 166,fig. 11a, b. Cretaceous. Essen, Ruhr Valley,Westphalia [Germany]. Polychaeta (fide Pilsbry &Sharp 1898: 246).typa, Throopella Greger, 1933: 373, 374, pl. 21. UpperDevonian, Cow Creek stage. Snyder Creek, centralMissouri, USA. Emerson (1962: 480) andStarobogatov (1974: 8) excluded this taxon fromthe Scaphopoda. Starobogatov (1974) placed it inthe toxeumorphorid Xenoconchia.undatum, Dentalium Defrance, 1819a: 72. Saint-Clément, near Angers [France]. Ditrupa (Polychaeta)(fide Pilsbry & Sharp 1898: 244).vayssierei, Dentalium Fenaux, 1942: 3, text-fig. 5.Recent, 80 m, off Carro, France (MediterraneanSea). Two syntypes (presumably in MOM).Polychaeta (fide V. Scarabino, in litt. 5.XI.2001).wilsoni, Dentalium Fraas, 1867a: 239, pl. 4, fig. 12;1867b: 95, pl. 1, fig. 12. Cretaceous. Marsaba,Palestine. Polychaeta (fide Pilsbry & Sharp 1898:246). Blanckenhorn (1927: 123, 124, pl. 1, figs 2-8) listed this as a junior synonym of Dentalium octocostatumFraas, 1867, which he treated as a validspecies of Dentalium, but he did not discuss thecontrary conclusion of Pilsbry & Sharp.AcknowledgementsThe first author is most indebted to P. Bouchet,B. Métivier (MNHN), S. Gofas (University ofMalaga), J. Taylor, K. Way, J. Pickering (BMNH)and D. Ivanov (MMSU) for making it possible towork at their institutions and for their helpwith the collections. I owe special thanks toV. Scarabino (Paris) for providing a wealth ofinformation and help.The second author thanks K. J. Boss (MCZ) andM. G. Harasewych (USNM) for allowing him touse their departmental libraries, T. R. Waller andJ. Thompson (USNM Paleobiology) for allowinghim to examine the fossil type material in theircollections, V. Héros (MNHN) for her help withthe Dautzenberg type localities, and R. I. Johnson(MCZ) for his fine hospitality and access to hispersonal library.The following colleagues provided us with valuableinformation on type material and synonyms, or sentus copies of obscure publications: A. Baldinger(MCZ), A. Beu (NZGS), R. Bieler (FMNH),E. Brunstad (Bergen, Norway), P. Callomon(Japan), K. Chinzei (FGK), G. Coan (CAS),A. Dhondt (KBINW), R. Eng (WSM), Y. Finet(MHNG), D. Geiger (SBMNH), B. Goddeeris(KBINW), D. Graf (ANSP), L. Groves (LACM),R. Janssen (SMF), E. Kools (CAS), S. Kosuge(IMT), K. B. Kvalsvik (ZMO), R. La Perna (Univ.Catania, Italy), A. Matsukuma (KyushuUniversity), R. Moolenbeek and A. N. Van derBijl (ZMA), G. Pastorino (Museo Argentino deCiencias Naturales), R. E. Petit (North MyrtleBeach, South Carolina), M. Recevik (MCZ),G. Rosenberg (ANSP), H. Saito (NSMT),K. Sindemark (SMNH), J. A. Sneli (TrondheimBiological Station, Norway), J. Stilwell (JCU),R. Symonds (UMZC), S. Takeuchi (SHM),F. Wesselingh (RMNH), and R. White (YPM).We most appreciate the reviews provided byPhilippe Bouchet and Victor Scarabino, whichenhanced the accuracy and presentation of thiscatalog. Helmut Zibrowius (Centre d’Océanologiede Marseille) provided helpful commentson various species that were originally describedas scaphopods but subsequently determined to benon-molluscan. Any remaining errors or omissionsare solely the authors’ responsibility.REFERENCESThe publication dates for books and articles publishedin journals has been verified, to the greatest extent possible,by reference to bound journal volumes and tocompilations of publication dates for certain journals.The publication dates for the following works arebased upon these sources: Chenu’s Illustrationsconchyliologiques (Sherborn & Smith 1911); Journal deconchyliologie (Fischer-Piette 1937; Winckworth1936); Manual of Conchology (Vanatta 1927); TheNautilus (Coan & Harasewych 1993); Proceedings(and Transactions) of the Zoological Society of London(Duncan 1937); Bulletin (and Proceedings) of theUnited States National Museum (Anonymous 1947,1971); Bullettino malacologico Italiano and Bullettinodella Società Malacologica Italiana (Manganelli &Cianfanelli 2001).ABBOTT R. T. 1974. — American Seashells. SecondEdition. Van Nostrand Reinhold Co., New York,663 p., 24 pls.ZOOSYSTEMA • 2004 • 26 (4)669
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