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LICHENS AND LICHEN. PARASITES

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218 B.A.N.Z. ANTARCTIC RESEAHCII ESPEDTTlON<br />

axils which are wider, somewhat inflated, fistnlose (perhaps from degeneration of old soredia rather<br />

than as in R. iqzjluta), holes long elliptic, irregularly placed, often along the edges of the branches,<br />

colour when fresh unlrnown (preserved specimens xiearly white, variously spotted with brown),<br />

true cortex lacking, replaced by mechanical tissue about 40p thiclr, highly gelified, of longitudinal<br />

branched hyphae with very narrow lumina; algal layer 20p thiclr, continuous, of closely packed<br />

cells 5-6p in diameter (absent in one branch sectioned except in soralia) ; central portion stuffed<br />

with thick-walled, browriish hyphae, 34p in diameter, very loosely woven; soralia elliptic, white,<br />

somewhat eroded, soredia of a few closely packed algal cells, 5-6p in diameter, more or less angular<br />

from mutual pressure, snrrounded by a few layers of very slender, thin-walled, hyaline hyphae,<br />

resting on a thin layer of the same type of hyphae. Apothecia and spermogoiiia not seen.<br />

Growing tangled with Cladoniaceae. This species belongs in the sect. Ecorticatae Stnr. and is<br />

closely related to R. injlata and R. genkztlatu, perhaps a sorediose coi~ditioli derived from one of<br />

them. Only more and better material can solve the probiem raised. As these species have often<br />

been confused and may later be found in our area, I have provided descriptions below, based on<br />

material in the herbarium of Thomas Taylor at IIarvard University.<br />

Macquarie Island, Featherbed Flat, Sta. Sla, R.A.N.Z.A.R.E. 531-25,531-26,531-27,531-28,<br />

531-29,531-30; north end of island, Sta. 81, B.A.N.Z.A.R.E. 540-14.<br />

Ranudinu linearis (Sw.) Ach., Lichenogr. ITniv., 598; 1810.<br />

Lichen li~benris (Sw.), Meth. Musc., 3G, pl. 2, fig. 3 ; 1781.<br />

Type : New Zealand, no collector cited. The following descriptior~<br />

is based on New Zealand,<br />

J. D. Hooker (Voy. "Erebus & Terror" in Taylor Herb. at Harvard Univ. sub R. gelziculata<br />

Hook. f. & Tayl., plants marltecl "c" by 3liiller-Argan and determined by him as R. linearis. (cf.<br />

Flora, 71, 132 ; 1888).<br />

Thallus 5 cm. tall, basal portion reticulate foraminous (as in R. reticliluta [Noehd.]<br />

IZrmplhbr.), repeatedly dichotomons, the hraiiches flattened, somewllat rlervose costate becoming<br />

canaliculate with the margins slightly inrolled, ultimate branches very narrow but still flat, apo-<br />

thecia borne along the margins on short stipes, not producing a bend of the frond at the point of<br />

attachment; probably whitish ashy when fresh, now light pinkish cinnamon; cortex lacking, being<br />

replaced by mechanical tissue which is 3510p thick (with thickenings up to 75p), highly gelified,<br />

of predominantly longitudinal, conglutinate, thick-wallecl hyphae about 7-8p in diameter, lumen<br />

about lp; algal layer 40-75p thick, sometimes completely filling the central portion,:tending to<br />

occur in subspherical colonies with spaces between the colonies filled with loosely woven medullary<br />

hyphae about 5p in diameter, lumen about lp; central portion with large air spaces but not truly<br />

hollow ; K-. c<br />

Apothecia up to 2.mm. in diameter, disc slightly coilcave at first with slightly elevated margin,<br />

becoming very convex and immarginate, evciple smooth, pale ochraceous buff from whitish pruina ;<br />

amphithecia1 cortex highly gelified, 9Op thick, thinning toward the margin, outer 35p a palisade,<br />

rest of irregularly periclinal hyphae, similar to the mechanical tissue of the thallns; al, we abnn-<br />

dant under the central portion of the thecium, thinning and disappearing under the outer two<br />

thirds, medullary hyphae not differentiated apart from the algal portion; hypothecium about 35p<br />

thick, of very slender, densely woven hyphae, thiniiing to about 2OP at the margin, hyphae increas-<br />

ingly periclinal and expanding upward to the margin of the thecinm as a parathecium; thecium<br />

about 75p tall; paraphyses very slender, dichotoinously branched above the asci, tips not conspicn-<br />

ously thickened, epithecium gelified with minute crystals; asci cylindric clavate, 8-spored,

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