waders and their estuarine food supplies - Vlaams Instituut voor de ...

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$Wetenschap - \Nijverheid- Handel$

15 Versatility of male curlews (Numenius arquata) preying upon Nereis diversicolor: deploying contrasting capture mode dependent on prey availability. 315 16 How Oystercatchers and Curlews successively deplete clams. 333 Samenvatting 345 References 367
INTRODUCTION
Page 1 and 2: WADERS AND THEIR ESTUARINE FOOD SUP
Page 3 and 4: plia^ohi. WADERS AND THEIR ESTUARI
Page 5 and 6: Waders and their estuarine food sup
Page 7 and 8: WADERS AND THEIR ESTUARINE FOOD SUP
Page 10 and 11: figuren: Dick Visser omslagfoto: Ja
Page 15 and 16: That science is making progress, ma
Page 17 and 18: INTRODUCTION The remnants of brushw
Page 19 and 20: When hcnlhic bivalves extend llien
Page 21 and 22: the burrow depths and on the fracti
Page 23 and 24: INTRODUCTION Ms,i invest 409 of (he
Page 25 and 26: able lo sw itch 10 oiher prey which
Page 27 and 28: Long-term, broadly-based research c
Page 29 and 30: Chapter 1 SEASONAL VARIATION IN BOD
Page 31 and 32: SEASONAL VARIATION IN BODY WEIGHT O
Page 33 and 34: Tahle 1 Weight loss ('•- ± SE) m
Page 39 and 40: i 60 50 40 30 20 10 0 • INFESTED
Page 41 and 42: 240- SEASONAL VARIATION IN BODY WEI
Page 43 and 44: 20 10 0 -10 -20h 2 3 4 5 6 7 8 9 se
Page 45 and 46: a E 400 - S. plana 35 mm 320 240 16
Page 49 and 50: Chapter 2 HOW THE FOOD SUPPLY HARVE
Page 51 and 52: FOOD SUPPLY HARVESTABLE BY WADERS H
Page 53 and 54: Methods The study sites were situat
Page 55 and 56: less than that of bivalves, partly
Page 57 and 58: I Fig. 3). Peak condition was reach
Page 59 and 60: total biomass since most of those t
Page 61 and 62: on the basis of the preceding and t
Page 63 and 64:
However, for obvious reasons, we ma
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prey. A decrease in the prey densit
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Tahle 2. The intake rate ol < lyste
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* ' % -. . ; a X - . - * • ^ •
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(Zwarts & Wanink 1989). In quiet we
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general law relating handling time
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nearly twice as much time (0.79 s).
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4 5 6 7 8 9 size class of Corophium
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and annually. Second, the lower siz
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Prey switching Waders feeding on ti
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autumn and spring, and the contrary
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where they switch to surface-living
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Chapter 3 BURYING DEPTH OF THE BENT
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DEPTH AND SIPHON CROPPING IN SCROBI
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I DEPTH AND SIPHON CROPPING IN SCRO
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DEPTH AND SIPHON CROPPING IN SCROBI
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Table 3. Three-Way analysis Of vari
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Chapter 4 SIPHON SIZE AND BURYING D
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15 20 size (mm) Fig. 3. Cerasioderm
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10 size (mm) SIPHON SIZE AND DEPTH
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o, 7 01 5 | * CL 5- SIPHON SIZE AND
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4 Q) Si •a a. 16- 20- A predator:
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prey risk for an animal al a depth
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Table 6. Size al which there is a m
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* ' 1 /-/ "».-^*«C
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face and so expose themselves to a
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surface in die containers was varie
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50 OOF 310.00 I 5.00 ^ 1.00 3=" 0.5
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Tahle 1. Macoma and Scrobicularia.
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siphon would not have to reach the
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known siphon weight and burying dep
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ACCESSIBLE PREY ARE OFTEN IN POOR C
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1 2 3 4 burying depth (cm) Kin. I.
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I—1 • • : : : : ! - ! -|-r 0
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Chapter 7 DOES AN OPTIMALLY FORAGIN
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OPTIMAL FORAGING AND THE FUNCTIONAL
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100 200 300 prey density (n-m-2) Fi
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Table I. Results of five one way an
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Table 2. Results of eight iwo-wav a
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prey density II III Fig. 12. Freque
Page 147 and 148:
Page 149:
PREY SIZE SELECTION AND INTAKE RATE
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Predicted 'passive size selection'
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lig. 2. Larger Mussels are less ava
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Fig. 5. Scrobicularia plana. Size c
Page 158 and 159:
and maiiv are too thick-shelled to
Page 160 and 161:
The measurements of handling time i
Page 162 and 163:
1 2 3 4 5 6 7 probing depth (cm PRE
Page 164 and 165:
valves may vary by a factor of two
Page 166 and 167:
optimal foraging model, the minimum
Page 168 and 169:
their gut is full? Do they reduce t
Page 171 and 172:
PREY PROFITABILITY AND INTAKE RATE
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catchers to take only certain prey
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licult error of estimate arose if p
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Counts of feeding and non-feeding b
Page 179 and 180:
Page 181 and 182:
20 30 40 50 shell length (mm) PREY
Page 183 and 184:
Page 185 and 186:
Nereis Arenicola tipuia earthwo'm M
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take rate. We might therefore expec
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prey species, using parallel slopes
Page 191 and 192:
5.0 4.0 1-3.0 a & • % * • * •
Page 193 and 194:
time during the feeding period. As
Page 195 and 196:
winter. Similarly, handling time in
Page 197 and 198:
5.0 - f-4.0 S 3.0 in I 20 a 2 a | 1
Page 199 and 200:
situation arrived in the western pa
Page 201 and 202:
Page 203 and 204:
• ' • 14 PREY PROFITABILITY AND
Page 205 and 206:
Notes lo appendix: PREY PROFITABILI
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Chapter 10 WHY OYSTERCATCHERS HAEMA
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2 4 6 8 10 time on feeding area (h)
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80- 60- o 80 60- 40- 20- INTAKE RAT
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est of bod\ behind: an estimated 22
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INTAKE RATE AND PROCESSING RATE IN
Page 218 and 219:
Wanink 1993). The energy content of
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(7) Age All studies dealt with adul
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irds over long periods. As an examp
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Discussion There is no difference i
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comparison between the weight of ih
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. 1', L ! •
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Introduction PREDICTING SEASONAL AN
Page 232 and 233:
PREDICTING SEASONAL AND ANNUAL FLUC
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Page 236 and 237:
Page 238 and 239:
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1000 r 1 II" 1 - r^Jan'80 PREDICTIN
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Page 244 and 245:
Page 246 and 247:
Page 248 and 249:
Page 250 and 251:
Page 252 and 253:
IOO BO 60 40 20 0 PREDICTING SEASON
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Page 256 and 257:
Page 258 and 259:
Page 260 and 261:
Page 262 and 263:
Page 265 and 266:
WHY KNOT TAKE MEDIUM-SIZED MACOMA W
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in Zwarts & Esselink 1989). The len
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shell length (mm) FIR. 3. Peringia.
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fused, specimens larger than this w
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50 100- s s I — f = S. plana, n=2
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area is twice as large as the touch
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10 15 lower size threshold (mm) Fig
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lime (Hughes 1979). This would furt
Page 281 and 282:
WHY KNOT TAKE MEDIUM-SIZED MACOMA T
Page 283 and 284:
Chapter 13 ANNUAL AND SEASONAL VARI
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VARIATION IN FOOD SUPPLY OF KNOT AN
Page 287 and 288:
Two sites (N and M in Fig. 2) were
Page 289 and 290:
20 30 VARIATION IN FOOD SUPPLY OF K
Page 291 and 292:
20 l i r'l ' i •o j^y^T n ^ ' " ^
Page 293 and 294:
July ' Aug ' Sept Fig. 9. Proportio
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available. There must, however, be
Page 297 and 298:
Chapter 14 SEASONAL TREND IN BURROW
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BURROWING AND FEEDING IN NEREIS SEA
Page 301 and 302:
Table 2. Results of a 2-way analysi
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June 1981 June 1982 is * N.MUD •
Page 305 and 306:
8 10 I 12 JZ 5. -g 1*» •e o i 16
Page 307 and 308:
6 - n=!080 5 • g 4 CP > i 3 CO CD
Page 309 and 310:
1985) and Curlew (/wails ,v, Lsseli
Page 311:
Chapter 15 VERSATILITY OF MALE CURL
Page 314 and 315:
Smid! 1951. Muus 1967, Wolff 1973,
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0.6 0.8 1.0 1.2 1.4 1.6 jaw lengih
Page 318 and 319:
Na oi both Nrieck Fig. 7. Numenius
Page 320 and 321:
6 8 10 12 14 16 18 worm length (cm)
Page 322 and 323:
too (Fig. 11 A). Indeed, the averag
Page 324 and 325:
VERSATILITY OF CURLEWS FEEDING ON N
Page 326 and 327:
total time spent al the surface. Th
Page 328 and 329:
2 4 . • 2.2 • 30 .-.2.0 to Q. 1
Page 331 and 332:
PREY DEPLETION BY OYSTERCATCHER AND
Page 333 and 334:
the rule: the observed lower limit
Page 335 and 336:
to locate the clams after they had
Page 337 and 338:
Curlew, by bury ing some hundreds o
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PREY DEPLETION BY OYSTERCATCHER AND
Page 341 and 342:
SAMENVATTING 345
Page 343 and 344:
Voorgeschiedenis In 1960 verscheen
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maken. Als gebied A een grotere voe
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omdat ze nog vrijwel niets wegen. l
Page 349 and 350:
kunnen opzuigen. of diep leven en z
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van de uitgerekte sifo over het wad
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het toen niet gemakkelijk hebben ge
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doende nonnetjes van 10 tot 15 mm l
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aangevoerde voedsel en de wulp past
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zijn dan twee jaar en wulpen geen s
Page 361 and 362:
was dan ook dank/ij de inzet van al
Page 363 and 364:
REFERENCES 367
Page 365 and 366:
Allen RL. 1983. Feeding behaviour o
Page 367 and 368:
BreyT. 1989. Der Einfluss physikali
Page 369 and 370:
latie tol chironoiiiiilen en de wai
Page 371 and 372:
huch der Vogel Milteleuropas, Band
Page 373 and 374:
llolmann II. & H. Huerschelmann 196
Page 375 and 376:
l-cndrem D.W. 1984. Flocking, feedi
Page 377 and 378:
Pekkarinen M. 1984. Regeneration of
Page 379 and 380:
lion of Mussels Mytilus edulis hy O
Page 381 and 382:
lulal Dal esiuanes: a comparison of
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