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WADERS AND THEIR ESTUARINE FOOD SUP
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plia^ohi. WADERS AND THEIR ESTUARI
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Waders and their estuarine food sup
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WADERS AND THEIR ESTUARINE FOOD SUP
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figuren: Dick Visser omslagfoto: Ja
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15 Versatility of male curlews (Num
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That science is making progress, ma
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INTRODUCTION The remnants of brushw
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When hcnlhic bivalves extend llien
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the burrow depths and on the fracti
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INTRODUCTION Ms,i invest 409 of (he
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able lo sw itch 10 oiher prey which
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Long-term, broadly-based research c
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Chapter 1 SEASONAL VARIATION IN BOD
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SEASONAL VARIATION IN BODY WEIGHT O
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Tahle 1 Weight loss ('•- ± SE) m
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SEASONAL VARIATION IN BODY WEIGHT O
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SEASONAL VARIATION IN BODY WEIGHT O
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i 60 50 40 30 20 10 0 • INFESTED
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240- SEASONAL VARIATION IN BODY WEI
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20 10 0 -10 -20h 2 3 4 5 6 7 8 9 se
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a E 400 - S. plana 35 mm 320 240 16
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SEASONAL VARIATION IN BODY WEIGHT O
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Chapter 2 HOW THE FOOD SUPPLY HARVE
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FOOD SUPPLY HARVESTABLE BY WADERS H
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Methods The study sites were situat
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less than that of bivalves, partly
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I Fig. 3). Peak condition was reach
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total biomass since most of those t
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on the basis of the preceding and t
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However, for obvious reasons, we ma
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prey. A decrease in the prey densit
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Tahle 2. The intake rate ol < lyste
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* ' % -. . ; a X - . - * • ^ •
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(Zwarts & Wanink 1989). In quiet we
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general law relating handling time
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nearly twice as much time (0.79 s).
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4 5 6 7 8 9 size class of Corophium
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and annually. Second, the lower siz
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Prey switching Waders feeding on ti
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autumn and spring, and the contrary
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where they switch to surface-living
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Chapter 3 BURYING DEPTH OF THE BENT
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DEPTH AND SIPHON CROPPING IN SCROBI
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I DEPTH AND SIPHON CROPPING IN SCRO
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DEPTH AND SIPHON CROPPING IN SCROBI
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Table 3. Three-Way analysis Of vari
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Chapter 4 SIPHON SIZE AND BURYING D
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15 20 size (mm) Fig. 3. Cerasioderm
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10 size (mm) SIPHON SIZE AND DEPTH
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o, 7 01 5 | * CL 5- SIPHON SIZE AND
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4 Q) Si •a a. 16- 20- A predator:
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prey risk for an animal al a depth
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Table 6. Size al which there is a m
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* ' 1 /-/ "».-^*«C
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face and so expose themselves to a
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surface in die containers was varie
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50 OOF 310.00 I 5.00 ^ 1.00 3=" 0.5
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Tahle 1. Macoma and Scrobicularia.
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siphon would not have to reach the
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known siphon weight and burying dep
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ACCESSIBLE PREY ARE OFTEN IN POOR C
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1 2 3 4 burying depth (cm) Kin. I.
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I—1 • • : : : : ! - ! -|-r 0
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Chapter 7 DOES AN OPTIMALLY FORAGIN
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OPTIMAL FORAGING AND THE FUNCTIONAL
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100 200 300 prey density (n-m-2) Fi
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Table I. Results of five one way an
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Table 2. Results of eight iwo-wav a
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OPTIMAL FORAGING AND THE FUNCTIONAL
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prey density II III Fig. 12. Freque
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OPTIMAL FORAGING AND THE FUNCTIONAL
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PREY SIZE SELECTION AND INTAKE RATE
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Predicted 'passive size selection'
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lig. 2. Larger Mussels are less ava
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Fig. 5. Scrobicularia plana. Size c
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and maiiv are too thick-shelled to
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The measurements of handling time i
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1 2 3 4 5 6 7 probing depth (cm PRE
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valves may vary by a factor of two
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optimal foraging model, the minimum
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their gut is full? Do they reduce t
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PREY PROFITABILITY AND INTAKE RATE
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catchers to take only certain prey
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licult error of estimate arose if p
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Counts of feeding and non-feeding b
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PREY PROFITABILITY AND INTAKE RATE
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20 30 40 50 shell length (mm) PREY
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PREY PROFITABILITY AND INTAKE RATE
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Nereis Arenicola tipuia earthwo'm M
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take rate. We might therefore expec
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prey species, using parallel slopes
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5.0 4.0 1-3.0 a & • % * • * •
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time during the feeding period. As
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winter. Similarly, handling time in
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5.0 - f-4.0 S 3.0 in I 20 a 2 a | 1
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situation arrived in the western pa
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PREY PROFITABILITY AND INTAKE RATE
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• ' • 14 PREY PROFITABILITY AND
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Notes lo appendix: PREY PROFITABILI
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Chapter 10 WHY OYSTERCATCHERS HAEMA
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2 4 6 8 10 time on feeding area (h)
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80- 60- o 80 60- 40- 20- INTAKE RAT
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est of bod\ behind: an estimated 22
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INTAKE RATE AND PROCESSING RATE IN
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Wanink 1993). The energy content of
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(7) Age All studies dealt with adul
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irds over long periods. As an examp
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Discussion There is no difference i
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comparison between the weight of ih
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. 1', L ! •
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Introduction PREDICTING SEASONAL AN
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PREDICTING SEASONAL AND ANNUAL FLUC
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PREDICTING SEASONAL AND ANNUAL FLUC
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PREDICTING SEASONAL AND ANNUAL FLUC
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PREDICTING SEASONAL AND ANNUAL FLUC
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1000 r 1 II" 1 - r^Jan'80 PREDICTIN
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PREDICTING SEASONAL AND ANNUAL FLUC
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PREDICTING SEASONAL AND ANNUAL FLUC
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PREDICTING SEASONAL AND ANNUAL FLUC
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PREDICTING SEASONAL AND ANNUAL FLUC
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PREDICTING SEASONAL AND ANNUAL FLUC
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IOO BO 60 40 20 0 PREDICTING SEASON
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PREDICTING SEASONAL AND ANNUAL FLUC
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PREDICTING SEASONAL AND ANNUAL FLUC
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PREDICTING SEASONAL AND ANNUAL FLUC
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PREDICTING SEASONAL AND ANNUAL FLUC
- Page 262 and 263: PREDICTING SEASONAL AND ANNUAL FLUC
- Page 265 and 266: WHY KNOT TAKE MEDIUM-SIZED MACOMA W
- Page 267 and 268: in Zwarts & Esselink 1989). The len
- Page 269 and 270: shell length (mm) FIR. 3. Peringia.
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- Page 275 and 276: area is twice as large as the touch
- Page 277 and 278: 10 15 lower size threshold (mm) Fig
- Page 279 and 280: lime (Hughes 1979). This would furt
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- Page 283 and 284: Chapter 13 ANNUAL AND SEASONAL VARI
- Page 285 and 286: VARIATION IN FOOD SUPPLY OF KNOT AN
- Page 287 and 288: Two sites (N and M in Fig. 2) were
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- Page 293 and 294: July ' Aug ' Sept Fig. 9. Proportio
- Page 295 and 296: available. There must, however, be
- Page 297 and 298: Chapter 14 SEASONAL TREND IN BURROW
- Page 299 and 300: BURROWING AND FEEDING IN NEREIS SEA
- Page 301 and 302: Table 2. Results of a 2-way analysi
- Page 303 and 304: June 1981 June 1982 is * N.MUD •
- Page 305 and 306: 8 10 I 12 JZ 5. -g 1*» •e o i 16
- Page 307 and 308: 6 - n=!080 5 • g 4 CP > i 3 CO CD
- Page 309 and 310: 1985) and Curlew (/wails ,v, Lsseli
- Page 311: Chapter 15 VERSATILITY OF MALE CURL
- Page 315 and 316: lahle- I. Numenius arquala. Search
- Page 317 and 318: Npcck* Nereis taken in a rapid, sin
- Page 319 and 320: 100 • f 10 * 0.1 0.01 - 8 8 0 VER
- Page 321 and 322: VERSATILITY OF CURLEWS FEEDING ON N
- Page 323 and 324: VERSATILITY OF CURLEWS FEEDING ON N
- Page 325 and 326: since the bird was also observed du
- Page 327 and 328: 4 6 8 10 12 worm length (cm) Fig. 1
- Page 329: Chapter 16 HOW OYSTERCATCHERS AND C
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- Page 334 and 335: PREY DEPLETION BY OYSTERCATCHER AND
- Page 336 and 337: 2 3 4 5 6 7 8 size of Mya arenaria
- Page 338 and 339: Curlew when ihey both feed on clams
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- Page 344 and 345: Krabben. garnalen en vissen eten. e
- Page 346 and 347: moeten zoeken. De zoektijd is gerel
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- Page 350 and 351: SCHOL, MAAR OOK GARNAAL EN STRANDKR
- Page 352 and 353: tijd om de snavel diep in de grond
- Page 354 and 355: Kanoetstrandlopers en hun voedselaa
- Page 356 and 357: om te eten: deze zijn buiten bereik
- Page 358 and 359: WAAROM KANOETSTRANDLOPERS, SCHOLEKS
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SAMENVATTING daarom blij dai de mee
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'-i£?*r3U *3a* t V,
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•ins 11.. I on a tidal Hat in the
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ulaiion dynamics of Mya arenaria an
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Esselink P & I. /warts 1989. Season
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I ustart 1.1). A I) Wtat, R.T. Clar
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coasts: spatial and temporal interc
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of pre-migralory fattening on the t
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fl.li. Helgolander Meeresunters. 30
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I'laiicliiliys flesus population in
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Zwarts I... A-M. Blomert. P. Spaak