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waders and their estuarine food supplies - Vlaams Instituut voor de ...

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VERSATILITY OF CURLEWS FEEDING ON NEREIS<br />

VERSATILITY OF MALE CURLEWS NUMENIUS ARQUATA<br />

PREYING UPON NEREIS DIVERSICOLOR: DEPLOYING<br />

CONTRASTING CAPTURE MODES DEPENDENT<br />

ON PREY AVAILABILITY<br />

Curlews Numenius arquaia use 2 methods to capture Ragworms Nereis diversicolor. they search for<br />

worms at the surface, which are taken with a single peck (called N k) <strong>and</strong>/or they search for visual cues,<br />

such as burrow entrances, <strong>and</strong> probe <strong>de</strong>ep to extract the worm from the burrow (called Npmbc). It is argued<br />

that male Curlews select predominantly active worms, which are either grazing at the surface<br />

(N k) or are filter feeding within <strong>their</strong> burrows (Nprilbc <strong>and</strong> sometimes N k). Profitability (mg s" 1 h<strong>and</strong>ling<br />

i increases with worm length <strong>and</strong> is greater for N k. because of its shorter h<strong>and</strong>ling time, than for<br />

Nprobc. Worms of both prey types < c. 6 cm are ignored, since profitability is below the intake rate<br />

(mg s" 1 feeding). During low water there is a shift from Nprobc to Np

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